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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Cucujoidea

Familia: Boganiidae
Subfamiliae: Boganiinae - Paracucujinae
Name

Boganiidae Sen Gupta & Crowson, 1966

Type genus: Boganium Sen Gupta & Crowson, 1966 [stem = Bogani-]
References

Crowson, R.A. 1990: A new genus of Boganiidae (Coleoptera) from Australia, with observations on glandular openings, cycad associations and geographical distribution in the family. Journal of the Australian Entomological Society, 29: 91–99. DOI: 10.1111/j.1440-6055.1990.tb00324.x
Endrödy-Younga, S. 1991: Boganiidae (Coleoptera: Cucujoidea) associated with cycads in South Africa: two new species and a new synonym. Annals of the Transvaal Museum, 35: 285–293. ISSN: 0041-1752
Endrödy-Younga, S.; Crowson, R.A. 1986: Boganiidae, a new beetle family for the African fauna (Coleoptera: Cucujoidea). Annals of the Transvaal Museum, 34: 253–273. ISSN: 0041-1752
Lawrence, J.F.; Ślipiński, A. 2010: 10.1. Boganiidae Sen Gupta and Crowson, 1966. Pp. 282-286 in: Leschen, R.A.B.; Beutel, R.G.; Lawrence, J.F. (volume eds.) Coleoptera, beetles. Volume 2: Morphology and systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter. ISBN 3110190753 ISBN 9783110190755
Sen Gupta, T. & Crowson, R.A. 1966. A new family of cucujoid beetles, based on six Australian and one New Zealand genera. Annals and Magazine of Natural History (13) 9(97-99): 61-85. DOI: 10.1080/00222936608651639 Paywall. Reference page.
Sen Gupta, T. & Crowson, R.A. 1969. Further observations on the family Boganiidae, with definition of two new families Cavognathidae and Phloeostichidae. Journal of Natural History 3(4): 571–590. DOI: 10.1080/00222936900770481 Paywall. Reference page.

Links

Boganiidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD

Boganiidae is a family of beetles, in the suborder Polyphaga.[1][2] Members of the family are found in southern Africa, Australia and New Caledonia. Adults and larvae are pollenivorous, feeding on the pollen of cycads and Myrtaceae, Meliaceae, Cunoniaceae and Elaeocarpaceae angiosperms.[3] Metacucujus and Paracucujus act as pollinators for cycads Encephalartos and Macrozamia respectively. This association with cycads goes back to at least the Mid-Cretaceous, with an extinct form being found with preserved cycad pollen in 99 million year old Burmese amber.[4]
Subfamilies and genera

†Palaeoboganium Liu et al. 2017[5] Daohugou, China, Callovian
Subfamily Boganiinae Sen Gupta and Crowson 1966
Afroboganium Endrödy-Younga & Crowson, 1986 Africa
Boganium Sen Gupta & Crowson, 1966 Australia
†Cretoboganium Cai and Huang 2018[6] Burmese amber, Myanmar, Cenomanian
Paracucujinae Endrödy-Younga & Crowson, 1986
Athertonium Crowson, 1990 Australia
†Cretoparacucujus Cai & Escalona, 2018[4] Burmese amber, Myanmar, Cenomanian
Dzumacium Escalona, Lawrence, Wanat & Ślipiński, 2015 New Caledonia
Metacucujus Endrödy-Younga & Crowson, 1986 Africa
Paracucujus Sen Gupta & Crowson, 1966 Australia

References

"Archived copy". Archived from the original on 2012-10-03. Retrieved 2009-11-21.
"Cucujoidea - Nomen.at - animals and plants".
Escalona, Hermes E.; Lawrence, John F.; Wanat, Marek; Ślipiński, Adam (July 2015). "Phylogeny and placement of Boganiidae (Coleoptera, Cucujoidea) with a review of Australian and New Caledonian taxa: Phylogeny of Boganiidae". Systematic Entomology. 40 (3): 628–651. doi:10.1111/syen.12126.
Cai, Chenyang; Escalona, Hermes E.; Li, Liqin; Yin, Ziwei; Huang, Diying; Engel, Michael S. (September 2018). "Beetle Pollination of Cycads in the Mesozoic". Current Biology. 28 (17): 2806–2812.e1. doi:10.1016/j.cub.2018.06.036.
Liu, Zhenhua; Ślipiński, Adam; Lawrence, John F.; Ren, Dong; Pang, Hong (2018-03-16). "Palaeoboganium gen. nov. from the Middle Jurassic of China (Coleoptera: Cucujoidea: Boganiidae): the first cycad pollinators?". Journal of Systematic Palaeontology. 16 (4): 351–360. doi:10.1080/14772019.2017.1304459. ISSN 1477-2019.
Cai, Chenyang; Huang, Diying (February 2019). "First boganiine beetle in mid-Cretaceous amber from northern Myanmar (Coleoptera: Boganiidae)". Proceedings of the Geologists' Association. 130 (1): 81–86. doi:10.1016/j.pgeola.2018.09.004. hdl:1983/aa67ee65-98bb-475c-9b23-5fb03e8715d6.

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