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dus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Cladus: Pterygota
Cladus: Metapterygota
Cladus: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Ordo: Diptera
Subordo: Brachycera
Infraordo: Asilomorpha
Superfamilia: Asiloidea
Familia: Bombyliidae
Subfamiliae: Anthracinae - Bombyliinae - Crocidiinae - Ecliminae - Gerontinae - Lomatiinae - Phthiriinae - Sysstropinae - Toxophorinae - Usiinae

Overview of genera (230)

Acreophthiria - Acreotrichus - Acrophthalmyda - Adelidea - Adelogenys - Aldrichia - Aleucosia - Alomatia - Amictus - Amphicosmus - Anastoechus - Anisotamia - Anthrax - Antonia - Antoniaustralia - Apatomyza - Aphoebantus - Apiformyia - Apolysis - Astrophanes - Atrichochira - Australiphthiria - Australoechus - Balaana - Beckerellus - Bombomyia - Bombylella - Bombylisoma - Bombylius - Brachyanax - Brachydemia - Bromoglycis - Brychosoma - Bryodemina - Cacoplox - Caecanthrax - Callostoma - Callynthrophora - Canariellum - Chalcochiton - Choristus - Chrysanthrax - Colossoptera - Comptosia - Conomyza - Cononedys - Conophorina - Conophorus - Corsomyza - Coryprosopa - Crocidium - Cryomyia - Cyananthrax - Cyllenia - Cyrtomyia - Cytherea - Cyx - Dasypalpus - Desmatomyia - Desmatoneura - Deusopora - Diatropomma - Dicranoclista - Diochanthrax - Dipalta - Diplocampta - Dischistus - Docidomyia - Doddosia - Dolichomyia - Doliogethes - Eclimus - Edmundiella - Efflatounia - Enica - Epacmoides - Epacmus - Eremyia - Eristalopsis - Eucessia - Euchariomyia - Euprepina - Eurycarenus - Euryphthiria - Eusurbus - Exechohypopion - Exepacmus - Exhyalanthrax - Exoprosopa - Exoprosopini - Gazlia - Geminaria - Geron - Gnumyia - Gonarthrus - Gyrocraspedum - Hallidia - Hemipenthes - Heteralonia - Heterostylum - Heterotropus - Hyperalonia - Hyperusia - Inyo - Isocnemus - Kapu - Karakumia - Laminanthrax - Larrpana - Laurella - Legnotomyia - Lepidanthrax - Lepidochlanus - Lepidophora - Ligyra - Litorhina - Lomatia - Lordotus - Macrocondyla - Mallophthiria - Mancia - Mandella - Mariobezzia - Marleyimyia - Megapalpus - Megaphthiria - Meomyia - Merleus - Metacosmus - Micomitra - Microrhina - Munjua - Muwarna - Myonema - Neacreotrichus - Nectaropota - Neobombylodes - Neodiplocampta - Neodischistus - Neosardus - Nomalonia - Nothoschistus - Notolegnotus - Notolomatia - Oestranthrax - Oestrimyza - Ogcodocera - Oligodranes - Oncodosia - Oniromyia - Othniomyia - Pachyanthrax - Palintonus - Palirika - Pantarbes - Pantostomus - Parachistus - Paracosmus - Paradiplocampta - Parageron - Paranthrax - Parasystoechus - Paratoxophora - Paravilla - Parisus - Peringueyimyia - Petrorossia - Phthiria - Phthiriinae - Pilosia - Pipunculopsis - Platamomyia - Plesiocera - Poecilanthrax - Poecilognathus - Prorachthes - Prorostoma - Prothaplocnemis - Pseudopenthes - Pteraulacodes - Pteraulax - Pterobates - Pusilla - Pygocona - Relictiphthiria - Rhynchanthrax - Satyramoeba - Semiramis - Sericosoma - Sericothrix - Sericusia - Sinaia - Sisyromyia - Sisyrophanus - Sosiomyia - Sparnopolius - Sphenoidoptera - Spogostylum - Staurostichus - Stomylomyia - Stonyx - Synthesia - Systoechus - Systropus - Thevenetimyia - Thraxan - Thyridanthrax - Tillyardomyia - Timiomyia - Tmemophlebia - Tomomyza - Tovlinius - Toxophora - Triplasius - Triploechus - Turkmeniella - Usia - Veribubo - Villa - Villoestrus - Walkeromyia - Wurda - Xenoprosopa - Xenox - Xerachistus - Xeramoeba - Ylasoia - Zaclava - Zinnomyia - Zyxmyia

Name

Bombyliidae Latreille, 1802

Vernacular names
English: Bee fly
한국어: 재니등에
日本語: ツリアブ科
Polski: bujankowate
Suomi: Kimalaiskärpäset
Türkçe: Arı sineğigiller

Bombyliidae is a large family of flies with hundreds of genera, although their life cycles are not well known. Adults generally feed on nectar and pollen, thus are pollinators of flowers. They superficially resemble bees, thus are commonly called bee flies, and this may offer the adults some protection from predators. In parts of East Anglia locals refer to them as 'beewhals', thanks to their tusk-like appendages.

The larval stages are predators or parasitoids of other insect eggs and larvae. The adult females usually deposit eggs in the vicinity of possible hosts, quite often in the burrows of beetles or wasps/solitary bees. Where most often in the insect world parasitoids are highly specific in the host species that they will infect, some bombyliids are opportunistic and will use a variety of hosts.

While bombyliids have a great variety of species, rarely are individuals of any one species abundant, and this is perhaps one of the poorest known families of insects relative to its size. There are at least 4,500 described species, and probably thousands as of yet undescribed.

Species include:

* Major bee-fly (Bombylius major)
* Williston's bee-fly (Poecilanthrax willistoni)
* Alkali bee fly (Heterostylum robustum)

Genera

* Acanthogeron Bezzi, 1925
* Acreophthiria Evenhuis, 1986
* Acreotrichus Macquart, 1840
* Acrophthalmyda Bigot, 1858
* Adelidea Macquart, 1840
* Adelogenys Hesse, 1938
* Aldrichia Coquillett, 1894
* Alepidophora Cockerell, 1909
* Aleucosia Edwards, 1934
* Alomatia Cockerell, 1914
* Amictites Hennig, 1966
* Amictus Wiedemann, 1817
* Amphicosmus Coquillett, 1891
* Anastoechus Osten-Sacken, 1877
* Anisotamia Macquart, 1840
* Anthrax Scopoli, 1763
* Antonia Loew, 1856
* Antoniaustralia Becker, 1913
* Apatomyza Wiedemann, 1820
* Aphoebantus Loew, 1872
* Apolysis Loew, 1860
* Astrophanes Osten Sacken, 1877
* Atrichochira Hesse, 1956
* Australiphthiria Evenhuis, 1986
* Australoechus Greathead, 1995
* Balaana Lambkin & Yeates, 2003
* Beckerellus Greathead, 1995
* Bombomyia Greathead, 1995
* Bombylella Greathead, 1995
* Bombylisoma Rondani, 1856
* Bombylius Linnaeus, 1758
* Brachyanax Evenhuis, 1981
* Brachydemia Hull, 1973
* Bromoglycis Hull, 1971
* Brychosoma Hull, 1973
* Bryodemina Hull, 1973
* Cacoplox Hull, 1970
* Caecanthrax Greathead, 1981
* Callostoma Macquart, 1840
* Callynthrophora Schiner, 1868
* Canariellum Strand, 1928
* Chalcochiton Loew, 1844
* Choristus Walker, 1852
* Chrysanthrax Osten Sacken, 1886
* Colossoptera Hull, 1973
* Comptosia Macquart, 1840
* Conomyza Hesse, 1956
* Cononedys Hermann, 1907
* Conophorina Becker, 1920
* Conophorus Meigen, 1803
* Corsomyza Wiedemann, 1820
* Coryprosopa Hesse, 1956
* Crocidium Loew, 1860
* Cryomyia Hull, 1973
* Cyananthrax Painter, 1959
* Cyllenia Latreille, 1802
* Cyrtomyia Bigot, 1892
* Cytherea Fabricius, 1794
* Cyx Evenhuis, 1993
* Dasypalpus Macquart, 1840
* Desmatomyia Williston, 1895
* Desmatoneura Williston, 1895
* Deusopora Hull, 1971
* Diatropomma Bowden, 1962
* Dicranoclista Bezzi, 1924
* Diochanthrax Hall, 1975
* Dipalta Osten Sacken, 1877
* Diplocampta Schiner, 1868
* Dischistus Loew, 1855
* Docidomyia White, 1916
* Doddosia Edwards, 1934
* Dolichomyia Wiedemann, 1830
* Doliogethys Hesse, 1938
* Eclimus Loew, 1844
* Edmundiella Becker, 1915
* Efflatounia Bezzi, 1925
* Enica Macquart, 1834
* Epacmoides Hesse, 1956
* Epacmus Osten Sacken, 1886
* Eremyia Greathead, 1996
* Eristalopsis Evenhuis, 1985
* Eucessia Coquillett, 1886
* Euchariomyia Bigot, 1888
* Euprepina Hull, 1971
* Eurycarenus Loew, 1860
* Euryphthiria Evenhuis, 1986
* Eusurbus Roberts, 1929
* Exechohypopion Evenhuis, 1991
* Exepacmus Coquillett, 1894
* Exhyalanthrax Becker, 1916
* Exoprosopa Macquart, 1840
* Geminaria Coquillett, 1894
* Geron Meigen, 1820
* Glaesamictus Hennig, 1966
* Gnumyia Bezzi, 1921
* Gonarthrus Bezzi, 1921
* Gyrocraspedum Becker, 1913
* Hallidia Hull, 1970
* Hemipenthes Loew, 1869
* Heteralonia Rondani, 1863
* Heterostylum Macquart, 1848
* Heterotropus Loew, 1873
* Hyperalonia Rondani, 1863
* Hyperusia Bezzi, 1921
* Inyo Hall & Evenhuis, 1987
* Isocnemus Bezzi, 1924
* Kapu Lambkin & Yeates, 2003
* Karakumia Paramonov, 1927
* Laminanthrax Greathead, 1967
* Larrpana Lambkin & Yeates, 2003
* Laurella Hull, 1971
* Legnotomyia Bezzi, 1902
* Lepidanthrax Osten Sacken, 1886
* Lepidochlanus Hesse, 1938
* Lepidophora Westwood, 1835
* Ligyra Newman, 1841
* Litorhina Bowden, 1975
* Lomatia Meigen, 1822
* Lordotus Loew, 1863
* Macrocondyla Rondani, 1863
* Mallophthiria Edwards, 1930
* Mancia Coquillett, 1886
* Mandella[disambiguation needed] Evenhuis, 1983
* Mariobezzia Becker, 1913
* Marleyimyia Hesse, 1956
* Marmosoma White, 1916
* Megapalpus Macquart, 1834
* Megaphthiria Hall, 1976
* Melanderella Cockerell, 1909
* Meomyia Evenhuis, 1983
* Metacosmus Coquillett, 1891
* Micomitra Bowden, 1964
* Munjua Lambkin & Yeates, 2003
* Muscatheres Evenhuis, 1986
* Muwarna Lambkin & Yeates, 2003
* Myonema Roberts, 1929
* Neacreotrichus Cockerell, 1917
* Nectaropota Philippi, 1865
* Neobombylodes Evenhuis, 1978
* Neodiplocampta Curran, 1934
* Neodischistus Painter, 1933
* Neosardus Roberts, 1929
* Nomalonia Rondani, 1863
* Nothoschistus Bowden, 1985
* Notolomatia Greathead, 1998
* Oestranthrax Bezzi, 1921
* Oestrimyza Hull, 1973
* Ogcodocera Macquart, 1840
* Oligodranes Loew, 1844
* Oncodosia Edwards, 1937
* Oniromyia Bezzi, 1921
* Othniomyia Hesse, 1938
* Pachyanthrax François, 1964
* Pachysystropus Cockerell, 1909
* Palaeoamictus Meunier, 1916
* Palaeogeron Meunier, 1915
* Palintonus François, 1964
* Palrika Lambkin & Yeates, 2003
* Pantarbes Osten Sacken, 1877
* Pantostomus Bezzi, 1921
* Paracorsomyza Hennig, 1966
* Paradiplocampta Hall, 1975
* Parachistus Greathead, 1980
* Paracosmus Osten Sacken, 1877
* Parageron Paramonov, 1929
* Paranthrax Bigot, 1876
* Parasysteochus Hall, 1976
* Paratoxophora Engel, 1936
* Paravilla Painter, 1933
* Parisus Walker, 1852
* Perengueyimyia Bigot, 1886
* Petrorossia Bezzi, 1908
* Phthiria Meigen, 1803
* Pilosia Hull, 1973
* Pipunculopsis Bezzi, 1925
* Platamomyia Brèthes, 1925
* Plesiocera Macquart, 1840
* Poecilanthrax Osten Sacken, 1886
* Poecilognathus Jaennicke, 1867
* Praecytherea Théobald, 1937
* Prorachthes Loew, 1868
* Prorostoma Hesse, 1956
* Prothaplocnemis Bezzi, 1925
* Pseudopenthes Roberts, 1928
* Pteraulacodes Hesse, 1956
* Pteraulax Bezzi, 1921
* Pterobates Bezzi, 1921
* Pusilla Paramonov, 1954
* Pygocona Hull, 1973
* Relictiphthiria Evenhuis, 1986
* Rhynchanthrax Painter, 1933
* Satyramoeba Sack, 1909
* Semiramis Becker, 1913
* Semistoechus Hall, 1976
* Sericosoma Macquart, 1850
* Sericothrix Hall, 1976
* Sericusia Edwards, 1937
* Sinaia Becker, 1916
* Sisyromyia White, 1916
* Sisyrophanus Karsch, 1886
* Sosiomyia Bezzi, 1921
* Sparnopolius Loew, 1855
* Sphenoidoptera Williston, 1901
* Spogostylum Macquart, 1840
* Staurostichus Hull, 1973
* Stomylomyia Bigot, 1888
* Stonyx Osten Sacken, 1886
* Synthesia Bezzi, 1921
* Systoechus Loew, 1855
* Systropus Wiedemann, 1820
* Thevenetimyia Bigot, 1875
* Thraxan Yeates & Lambkin, 1998
* Thyridanthrax Osten Sacken, 1886
* Tillyardomyia Tonnoir, 1927
* Timiomyia Evenhuis, 1978
* Tithonomyia Evenhuis, 1984
* Tmemophlebia Evenhuis, 1986
* Tomomyza Wiedemann, 1820
* Tovlinius Zaitzev, 1979
* Toxophora Meigen, 1803
* Triplasius Loew, 1855
* Triploechus Edwards, 1937
* Turkmeniella Paramonov, 1940
* Usia Latreille, 1802
* Veribubo Evenhuis, 1978
* Verrallites Cockerell, 1913
* Villa Lioy, 1864
* Villoestrus Paramonov, 1931
* Walkeromyia Paramonov, 1934
* Wurda Lambkin & Yeates, 2003
* Xenoprosopa Hesse, 1956
* Xenox Evenhuis, 1984
* Xerachistus Greathead, 1995
* Xeramoeba Hesse, 1956
* Ylasoia Speiser, 1920
* Zaclava Hull, 1973
* Zinnomyia Hesse, 1955
* Zyxmyia Bowden, 1960

References
Bombyliid fly on Bidens laevis

* Hull, F.M. ,1973. Bee flies of the world. The genera of the family Bombyliidae.Washington (Smithsonian Institution Press) 687 pp. Keys subfamilies, genera (many generic placements superseded by Evenhuis & Greathead, 1999).
* Evenhuis, N.L. 1991.Catalog of genus-group names of bee flies (Diptera: Bombyliidae) Bishop Museum Bulletin of Entomology5: 1–105.
* Evenhuis, N.L. & Greathead, D.J. 1999. World catalog of bee flies (Diptera: Bombyliidae). Backhuys Publishers, Leiden, 756 pp.
* Bowden, J.,1980 Family Bombyliidae. pp. 381–430. In R.W. Crosskey (ed.), Catalogue of the Diptera of the Afrotropical Region, 1437 pp., London: British Museum (Natural History)
* Engel, E.O., 1932-1937. Bombyliidae. In: Die Fliegen der paläarktischen Region 4(3) ( Erwin Lindner, ed.): 1-619, pl. 1-15. E. Schweizerbart, Stuttgart.). Old and outdated, not easy to get and expensive but some of the only keys to taxa in the Palaearctic Region.
* Greathead & Evenhuis (Greathead, D.J., & N.L. Evenhuis, 1997. Family Bombyliidae. In: Contributions to a manual of Palaearctic Diptera Volume 2 (L. Papp & B. Darvas, eds.): 487-512. Science Herald, Budapest.) provide a key to the Palaearctic genera and (may) give references to available generic revisions.

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