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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Cladus: Macroheterocera
Superfamilia: Noctuoidea

Familia: Erebidae
Subfamilia: Calpinae
Tribus: Calpini - Ophiderini - Phyllodini

[list of tribus after Lafontaine & Fibiger (2006)]
Name

Calpinae Boisduval, 1840
Overview of assigned genera

Arabriga – Brana – Calyptra – Deinypena – Episparis – Eudocima – Ferenta – Gloriana – Goniocraspedon – Gonodonta – Graphigona – Hemiceratoides – Lepidodes – Lobophyllodes – Miniodes – Miniophyllodes – Oporophylla – Oraesia – Papuacola – Parafodina – Phyllodes – Plusiodonta – Thaidenakaoga – Wolframmeyia – Xylophylla
Overview of genera not assigned to a tribus

Cabralia – Caryonopera – Chibidokuga – Egnasia – Epharmottomena – Eudrapa – Hyperlopha – Hypsiforma – Leucanitis – Marsipiophora – Megacephalomana – Megacephalon – Melanephia – Meyrickella – Oglasa – Pachyplastis – Prionophora (Meyrick) – Pseudotolna – Synalissa – Tamba
References

Inoue, H. 1979: A new species of the Noctuidae from Japan and Korea. Tinea 10(28): 285–289. Reference page.
Kobayashi, H., 2009: A new genus and a new species of Calpinae from Thailand (Noctuidae, Lepidoptera). Tinea 21(1): 6–8.
Lafontaine, J.D. & Fibiger, M. 2006. Revised higher classification of the Noctuoidea (Lepidoptera). Canadian entomologist 138(5): 610–635. DOI: 10.4039/n06-012 Reference page.
Lafontaine, J.D. & Schmidt, B.C. 2010. Annotated check list of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico. Zookeys 40: 1–239. DOI: 10.3897/zookeys.40.414 Reference page.
Poole, R.W., 1989: Lepidopterorum Catalogus New Series Noctuidae Part 1. Fascicle 118 : 3–500.
Zahiri, R., Holloway, J.D., Kitching, I.J., Lafontaine, J.D., Mutanen, M. & Wahlberg, N. 2011 (online) 2012 (print). Molecular phylogenetics of Erebidae (Lepidoptera, Noctuoidea). Systematic Entomology 37(1): 102–124. DOI: 10.1111/j.1365-3113.2011.00607.x Reference page.

External links

Phlogeny of Noctuidae: Calpinae
World Checklist of Tribe Calpini (Lepidoptera: Noctuidae: Calpinae) (PDF)

The Calpinae are a subfamily of moths in the family Erebidae described by Jean Baptiste Boisduval in 1840. This subfamily includes many species of moths that have a pointed and barbed proboscis adapted to piercing the skins of fruit to feed on juice, and in the case of the several Calyptra species of vampire moths, to piercing the skins of mammals to feed on blood.[1] The subfamily contains some large moths with wingspans longer than 5 cm (2 in).

Taxonomy

Recent phylogenetic studies have greatly revised this subfamily. The subfamily was previously classified within the Noctuidae, but the redefinition of that family has reclassified many of that family's subfamilies, including Calpinae, into the family Erebidae. The Calpinae are most closely related to a clade including the subfamilies Eulepidotinae and Hypocalinae, which are also among the Erebidae. The tribes Anomini and Scoliopterygini, previously included in the Calpinae, were found to be distantly related and were reclassified into a separate subfamily as the Scoliopteryginae.[2][3]
Tribes

The Calpinae consist of three monophyletic tribes.[3]

Calpini
Ophiderini
Phyllodini

Previous taxonomy

The status of the former composition of the Calpinae was somewhat disputed; it was sometimes merged into the Catocalinae. Most of the calpine genera were not further classified. The phylogenetic structure of this group was essentially unresolved, and in many cases it was even doubtful whether the genera were indeed correctly placed in this subfamily.

Tribe Calpini[4]

Africalpe Krüger, 1939
Calyptra Ochsenheimer, 1816
Eudocima Billberg, 1820
Ferenta Walker, 1858
Gonodonta Hübner, 1818
Graphigona Walker, 1858
Oraesia Boisduval & Guenée, 1852
Plusiodonta Boisduval & Guenée, 1852
Tetrisia Walker, 1867

Tribe Gonopterini

Scoliopteryx

Genera incertae sedis

Cecharismena Möschler, 1890
Culasta Moore, 1881
Euryschema Turner, 1925
Epicyrtica Turner, 1908
Goniapteryx Perty, 1833
Hemiceratoides Strand, 1911
Pharga Walker, 1863
Phyprosopus Grote, 1872
Psammathodoxa Dyar, 1921
Radara Walker, 1862

Genera provisionally placed here (incomplete list); includes taxa sometimes separated in Ophiderinae

Abriesa
Acanthoprora
Agamana
Alabama
Amphigonia
Anachrostis
Anoba
Anomis
Antiblemma
Anticarsia
Arcte
Armana
Arthisma
Arugisa
Ascalapha
Avitta
Azeta
Baniana
Belciana
Bematha
Bendisodes (Catocalinae: Ophiusina?)
Boryzops
Brachycyttara
Brachyona
Brana
Brevipecten
Brontypena
Bulia
Calymniops
Cephena
Chilkasa
Chodda
Cissusa
Concana
Condate
Conosema
Corcobara
Coxina (Catocalinae: Ophiusina?)
Crithote
Crypsiprora
Cultripalpa
Cyclodes
Daddala
Dahlia
Daona
Diascia
Diatenes
Diphthera
Dunira
Dyops
Eclipsea
Egnasia
Egnasides
Egone
Ephyrodes
Epidromia
Episparis
Episparina
Episparonia
Epitausa
Eporectis
Erebus (Catocalinae?)
Ericeia
Eubryopterella
Eudesmeola
Eulepidotis
Eurythmus
Facidina
Falana
Fodina
Forsebia
Gabara
Gloriana
Goniocarsia
Gracillina
Helia (Catocalinae: Ophiusina?)
Hemipsectra
Heteranassa (Catocalinae: Ophiusina?)
Hemeroblemma
Heterormista
Heterospila
Hirsutopalpis
Hopetounia
Hulodes
Hypocala
Hyperlopha
Hypoprora
Hyposemansis
Hypospila
Hypsoropha
Ianius
Isogona
Itomia (Catocalinae: Ophiusina?)
Kakopoda (Catocalinae: Ophiusina?)
Latebraria
Ledaea
Leptotroga
Lesmone (Catocalinae: Ophiusina?)
Lignicida
Lineopalpa
Litocala
Litoprosopus
Lophozancla
Lyncestoides Hacker & Saldaitis, 2010
Maguda
Malatrogia
Manbuta
Marapana
Masca
Massala
Matella
Mecodina
Melipotis (Catocalinae: Ophiusina?)
Meranda
Mesosciera
Metalectra
Metallata
Metaphoenia
Mimophisma (Catocalinae: Ophiusina?)
Neogabara
Neostichtis
Obrima
Oglasa
Olulis
Olyssa
Ophyx
Orodesma
Orthozancla
Ortopla
Ossonoba
Oxygonitis
Oxyodes
Palpidia
Pangrapta
Panopoda
Panula
Parachabora
Parapadna
Parolulis
Perciana
Perinaenia
Pilipectus
Phyllodes
Phytometra
Pleurona
Polydesma (Catocalinae: Ophiusina?)
Polydesmiola
Praxis
Prolyncestis
Propatria
Prorocopis
Pseudorgyia
Pseudogyrtona
Ramadasa
Raparna
Rema
Rhapsa
Rhesala
Rhesalides
Rhosologia
Rhynchodina
Saroba
Savara
Schistorhynx
Selenisa (Catocalinae: Ophiusina?)
Selenoperas
Spectrophysa
Sphingomorpha
Stenocarsia
Stenoprora
Sundwarda
Syllectra
Sypna
Sypnoides
Talmala
Tamba
Thiacidas
Thoracolophotos
Throana
Thyrostipa
Thysania
Tinolius
Tipasa
Tiruvaca
Tolpia
Tolpiodes
Toxonprucha (Catocalinae: Ophiusina?)
Trigonochrostia
Trisula
Tropidtamba
Tyrissa (Catocalinae: Ophiusina?)
Ugia
Veia
Vestura
Xanthanomis
Zaleops (Catocalinae: Ophiusina?)

References

Florida Museum of Natural History (ed.). "Blood-feeding and Fruit-feeding Moths of Calpini Tribe (Noctuidae), Their Phylogeny and Classification". Archived from the original on 11 April 2015. Retrieved 21 April 2015.
Lafontaine, Donald; Schmidt, Christian (March 19, 2010). "Annotated check list of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico". ZooKeys (40): 26. doi:10.3897/zookeys.40.414.
Zahiri, Reza; et al. (2011). "Molecular phylogenetics of Erebidae (Lepidoptera, Noctuoidea)". Systematic Entomology. 37: 102–124. doi:10.1111/j.1365-3113.2011.00607.x. S2CID 84249695.

J.M. Zaspel and M.A. Branham (2008). "World Checklist of Tribe Calpini (Lepidoptera: Noctuidae: Calpinae)". University of Nebraska, Lincoln. Archived from the original on December 20, 2014. Retrieved July 8, 2010.

External links
Savela, Markku (August 2, 2019). "Calpinae Boisduval, 1840". Lepidoptera and Some Other Life Forms. Retrieved November 21, 2019.

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