Carcharodini

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Hesperiidae
Subfamilia: Pyrginae
Tribus: Carcharodini
Subtribus: Carcharodina

Genera: Agyllia – Arteurotia – Austinus – Bolla – Burca – Carcharodus – Clytius – Conognathus – Cyclosemia – Ernsta – Favria – Gomalia – Gorgopas – Hesperopsis – Iliana – Incisus – Mictris – Mimia – Morvina – Muschampia – Myrinia – Nisoniades – Noctuana – Ocella – Pachyneuria – Pellicia – Perus – Pholisora – Polyctor – Sophista – Spialia – Staphylus – Tiana – Viola – Viuria – Windia – Xispia

Name

Carcharodini Verity, 1940
References

Austin, G.T. 2008. Hesperiidae of Rondônia, Brazil: a new genus and species of Carcharodini (Pyrginae). Bulletin of the Allyn Museum 158: 1–7. Reference page.
Cong, Q., Zhang, J., Shen, J.H. & Grishin, N.V. 2019. Fifty new genera of Hesperiidae (Lepidoptera). Insecta Mundi 0731: 1–56. web. Reference page.
Lamas, G. 2004. (ed.) Checklist: Part 4A. Hesperioidea - Papilionoidea. In Heppner, J.B. (ed.) Atlas of Neotropical Lepidoptera. Vol.5A, Pt.4A. Assn. for Tropical Lepidoptera/Scientific Publishers, Gainesville. 439pp. Reference page.
Steinhauser, S.R. 1989. Taxonomic notes and descriptions of new taxa in the neotropical Hesperiidae. Part I. Pyrginae. Bulletin of the Allyn Museum 127: 1–70. Reference page.
Verity, R., 1940-53. La Farfalle diurne d'Italia. Vols 1-5. 1 Gen/Hesperides; 2 Lycaenida; 3 Papilionidae, Pieridae; 4 Libytheina, Danaina, Nympalina (part); 5 Satyridae. Casa Editrice Marzocco. Reference page.
Zhang, J., Brockmann, E., Cong, Q., Shen, J.H. & Grishin, N.V. 2020. A genomic perspective on the taxonomy of the subtribe Carcharodina (Lepidoptera: Hesperiidae: Carcharodini). Zootaxa 4748(1): 182–194. DOI: 10.11646/zootaxa.4748.1.10. Paywall. Reference page.

The Carcharodini are a tribe in the skipper butterfly subfamily Pyrginae. They are a very diverse but quite plesiomorphic and inconspicuous group distributed throughout the tropics.[1]

Formerly, when only four tribes of Pyrginae were recognized, they were included in the Pyrgini, which at that time contained a massive number of genera. But the Pyrginae have since been reorganized to make them and their tribes monophyletic, leading most modern authors to treat the Carcharodini as distinct tribe. However, the old circumscription of the Pyrgini was by and large just as correct from a phylogenetic perspective.[1]

The tribe of the Pyrgini sensu lato most closely related to the Pyrgini sensu stricto are the strikingly different Achlyodidini. As many consider it desirable to treat this lineage as a distinct tribe, the Carcharodini naturally need to be considered distinct too.[1]
Genera

These genera belong to the tribe Carcharodini:[1][2][3]

Agyllia Grishin, 2020
Arteurotia Butler & H. Druce, 1872
Bolla Mabille, 1903
Burca Bell & Comstock, 1948
Carcharodus Hübner, [1819]
Conognathus Felder & Felder, 1862
Cyclosemia Mabille, 1878 (eyed skippers)
Ernsta Grishin, 2020
Favria Tutt, 1906
Fuscocimex Austin, 2008
Gomalia Moore, 1879 (marbled or African mallow skippers)
Gorgopas Godman & Salvin, 1894
Hesperopsis Dyar, 1905 (sootywings)
Iliana Bell, 1937
Jera Lindsey, 1925
Mictris Evans, 1955
Mimia Evans, 1953
Morvina Evans, 1953
Muschampia Tutt, 1906
Myrinia Evans, 1953
Nisoniades Hübner, 1819
Noctuana E. Bell, 1937
Ocella Evans, 1953
Pachyneuria Mabille, 1888
Pellicia Herrich-Schaffer, 1870
Pholisora Scudder, 1872 (sootywings)
Polyctor Evans, 1953
Sophista Plötz, 1879
Spialia Swinhoe, 1912 (grizzled skippers & sandmen)
Staphylus Godman & Salvin, 1896 (scallopwings)
Viola Evans, 1953 (Viola skippers)
Windia H. Freeman, 1969
Xispia Lindsey, 1925

Footnotes

Brower (2009)
Li, Wenlin; Cong, Qian; Shen, Jinhui; Zhang, Jing; et al. (2019). "Genomes of skipper butterflies reveal extensive convergence of wing patterns". Proceedings of the National Academy of Sciences. 116 (13). doi:10.1073/pnas.1821304116. PMC 6442542.

Zhang, Jing; Brockmann, Ernst; Cong, Qian; Shen, Jinhui; et al. (2020). "A genomic perspective on the taxonomy of the subtribe Carcharodina (Lepidoptera: Hesperiidae: Carcharodini)". Zootaxa. 4748 (1). doi:10.11646/zootaxa.4748.1.10.

References

Brower, Andrew V.Z. (2009): Tree of Life Web Project – Carcharodini. Version of 2009-JUN-14. Retrieved 2009-DEC-26.