Clambidae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Elateriformia
Superfamilia: Scirtoidea

Familia: Clambidae
Subfamiliae: Acalyptomerinae – Calyptomerinae – Clambinae
incertae sedis: †Ptisma
Overview of genera

Acalyptomerus – Calyptomerus – Clambus – Loricaster – Sphaerothorax – †Eoclambus – †Ptisma
Name

Clambidae Fischer von Waldheim, 1821
Synonyms

†Ptismidae Kirejtshuk & Azar in Kirejtshuk et al., 2016 [syn.: Cai et al., 2019]

References
Primary references

Cai, C., Lawrence, J.F., Yamamoto, S., Leschen, R.A.B., Newton, A.F., Ślipiński, A., Yin, Z., Huang, D. & Engel, M.S. 2019. Basal polyphagan beetles in mid-Cretaceous amber from Myanmar: biogeographic implications and long-term morphological stasis. Proceedings of the Royal Society (B) 286(1894): 20182175. DOI: 10.1098/rspb.2018.2175 Open access. Reference page.
Kirejtshuk, A.G., Chetverikov, P.E., Azar, D. & Kirejtshuk, P.A. 2016. Ptismidae fam. nov. (Coleoptera, Staphyliniformia) from the Lower Cretaceous Lebanese amber. Cretaceous Research 59: 201-213. DOI: 10.1016/j.cretres.2015.10.027 Paywall. PDF. [Corrigendum: 61: 275. DOI: 10.1016/j.cretres.2016.02.004 Open access.] Reference page.

Additional references

Crowson, R.A. 1979: Observations on Clambidae (Coleoptera), with descriptions of a new genus and species and of several larvae. Revue suisse de zoologie 86(3): 611–623. BHL Reference page.
Endrödy-Younga, S. 1965: Clambiden-Studien (Coleoptera). Annales historico-naturales Musei nationalis hungarici, 57: 259–265. ISSN: 0521-4726
Endrödy-Younga, S. 1974: A revision of the described Australian and New Zealand species of the family Clambidae (Coleoptera) with description of a new species. Records of the South Australian Museum, 17(1): 1–10. PDF
Endrödy-Younga, S. 1978: A revision of the Oriental Clambidae with references to Papuan and Hawaiian species (Coleoptera: Staphylinoidea). Pacific insects, 18: 67–84. PDF
Endrödy-Younga, S. 1981: The American species of the familia Clambidae (Coleoptera: Eucinetoidea). Entomologia generalis, 7(1): 33–67.
Endrödy-Younga, S. 1986: New species and new records for the Palaearctic and Oriental faunas of the family Clambidae (Coleoptera, Eucinetoidea). Revue Suisse de Zoologie, 93(1): 89–115.
Endrödy-Younga, S. 1990: Clambidae of New Zealand (Coleoptera: Eucinetoidea). New Zealand journal of zoology, 17: 119–135. BUGZ
Endrödy-Younga, S. 1990: A revision of the Australian Clambidae (Coleoptera: Eucinetoidea). Invertebrate taxonomy, 4: 247–280. DOI: 10.1071/IT9900247
Endrödy-Younga, S. 1993: Clambidae of the Ethiopian region (Coleoptera: Eucinetoidea). Annals of the Transvaal Museum, 36: 109–138. ISSN: 0041-1752
Endrödy-Younga, S. 1995: Clambidae of Sulawesi with reference to the biogeography of the Sunda Islands (Coleoptera, Eucinetoidea). Systematic entomology, 20: 1–16. DOI: 10.1111/j.1365-3113.1995.tb00079.x
Endrödy-Younga, S. 1998: The genus Clambus in Central and South America (Coleoptera: Eucinetoidea: Clambidae). Annals of the Transvaal Museum, 36: 373–403. ISSN: 0041-1752
Endrödy-Younga, S. 1999: Two new Clambus species from New Zealand (Coleoptera: Eucinetoidea: Clambidae). New Zealand journal of zoology, 26(1): 39–42. DOI: 10.1080/03014223.1999.9518175
Leschen, R.A.B. 2005: 15.3. Clambidae Fischer, 1821. Pp. 438-443 in: Beutel, R.G. & Leschen, R.A.B. (volume eds.) Coleoptera, Beetles. Volume 1: Morphology and systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter. ISBN 3110171309
Catalogue of Palearctic Coleoptera. Vol. 3 ed. I. Lobl, & A. Smetana, Apollo Books, Stenstrup, Denmark, 2006 ISBN 87-88757-59-5, p.314

Links

Australian Faunal Directory
Clambidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD
Tree of Life Web Project. 2007. Clambidae. Minute beetles. Version 07 May 2007 (temporary) in The Tree of Life Web Project
Atlas of family Clambidae of Russia
EoL

Vernacular names
Deutsch: Punktkäfer
English: minute beetles
lietuvių: Taškavabaliai
русский: Кругляки

Clambidae is a family of beetles. They are known commonly as the minute beetles[2] or the fringe-winged beetles.[3] They are found worldwide on every continent except Antarctica.[2]

These are tiny beetles with bodies measuring 0.7 to 2mm in length. They are flattened to convex in shape and some can roll into a ball. Some are hairless, while some are quite hairy or scaly.[4] The margins of the wings are lined with long hairs.[2]

Clambids commonly feed on fungi.[4]

The family is divided into 5 genera and about 70 described species.[2] The largest and most widespread genus is Clambus, which occurs around the world. The genus Sphaerothorax is found in Australia and New Zealand.[5] Acalyptomerus is circumtropical.[6] The latter two genera have undergone long term morphological stasis, with fossils identifiable to both genera being known from the 99 million year old Burmese amber. The oldest records of the group are the extinct genera Eoclambus and Ptisma from the Barremian aged Lebanese amber.[1]

Genera:

Acalyptomerus
Calyptomerus
Clambus
Loricaster
Sphaerothorax

References

Cai, Chenyang; Lawrence, John F.; Yamamoto, Shûhei; Leschen, Richard A. B.; Newton, Alfred F.; Ślipiński, Adam; Yin, Ziwei; Huang, Diying; Engel, Michael S. (2019-01-16). "Basal polyphagan beetles in mid-Cretaceous amber from Myanmar: biogeographic implications and long-term morphological stasis". Proceedings of the Royal Society B: Biological Sciences. 286 (1894): 20182175. doi:10.1098/rspb.2018.2175. ISSN 0962-8452. PMC 6367173. PMID 30963875.
Majka, C. G., & Langor, D. (2009). Clambidae (Coleoptera) of Atlantic Canada. Journal of the Acadian Entomological Society 5(7), 32-40.
Clambidae. Integrated Taxonomic Information System (ITIS).
Lawrence, J.F., et al. 2000 onwards. Clambidae. Elateriformia (Coleoptera): descriptions, illustrations, identification, and information retrieval for families and subfamilies. Version 9 October 2005.
Endrödy-Younga, S. (1990). Clambidae of New Zealand (Coleoptera: Eucinetoidea). New Zealand Journal of Zoology 17(1), 119-36.
Endrödy-Younga, S. (1998). Acalyptomerus Crowson: the circumtropical genus of the family Clambidae (Coleoptera: Clambidae). Koleopterologische Rundschau 68, 199-203.