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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Superfamilia: Tortricoidea

Familia: Tortricidae
Subfamilia: Tortricinae
Tribus: Cnephasiini
Genus: Cnephasia
Subgenera: C. (Anoplocnephasia) – C. (Cnephasia) – C. (Cnephasiella) – C. (Hypostephanuncia)
Overview of assigned species

C. abrasana – C. daedalea – C. delnoyana – C. etnana – C. hellenica – C. heringi – C. incertana
Overview of species not assigned to a sub-genus

C. albatana – C. alfacarana – C. alticola – C. alticolana – C. amseli – C. asbolodes – C. asiatica – C. asseclana – C. bizensis – C. chrysantheana – C. cinereipalpana – C. communana – C. conspersana – C. constantinana – C. cupressivorana – C. disforma – C. disparana – C. divisana – C. ecullyana – C. ergastularis – C. facetana – C. finita – C. fiorii – C. fragosana – C. fulturata – C. galeotis – C. genitalana – C. graecana – C. grandis – C. gueneana – C. heinemanni – C. hunzorum – C. incinerata – C. josebai – C. jozefi – C. kasyi – C. kenneli – C. klimeschi – C. korvaci – C. laetana – C. lineata – C. longana – C. macrostoma – C. margelanensis – C. mediocris – C. microstrigana – C. minutula – C. nesiotica – C. nigripunctana – C. nigrofasciana – C. nowickii – C. olearis – C. orientana – C. oxyacanthana – C. palestinensis – C. parnassicola – C. pasiuana – C. personatana – C. phalarocosma – C. quentini – C. razowskii – C. regifica – C. sedana – C. semibrunneata – C. stachi – C. stephensiana – C. syriella – C. taganista – C. taurominana – C. tianshanica – C. tofina – C. tolli – C. tremewani – C. tripolitana – C. tristrami – C. ussurica – C. venusta – C. virginana – C. wimmeri – C. zangheriana – C. zelleri – C. zernyi
Overview of species not assigned to Cnephasia

C. catastrepta – C. chlorocrossa
Name

Cnephasia Curtis, 1826

Type species: Phalaena logiana Clerck
Synonyms

Anoplocnephasia Réal, 1953
Brachycnephasia Réal, 1953
Chephasia Razowski, 1977 (misspelling of Cnephasia)
Cnephasianella Benander, 1950
Cnephasiella Adamczewski, 1936
Cnephosia Desmarest, 1857 (misspelling of Cnephasia)
Hypostephanuncia Réal, 1951
Hypostephanuntia Razowski, 1977 (misspelling of Hypostephanuncia)
Sciaphila (Tortricidae) Treitschke, 1829

References

Alipanah, H. 2019. An overview of the tribe Cnephasiini (Lepidoptera: Tortricidae: Tortricinae) of Iran with description of a new species. Zootaxa 4661(3): 501–521. DOI: 10.11646/zootaxa.4661.3.5. Reference page.
Brown, J.W. 2005. World Catalogue of Insects. Volume 5: Tortricidae (Lepidoptera) 1–741. Apollo Books. Reference page.
Gastón, J., Vives Moreno, A. & Revilla, T. 2017. Descripción de tres especies nuevas de la Familia Tortricidae en la Península Ibérica (Lepidoptera: Tortricidae). SHILAP Revista de Lepidopterología 45(180): 689-698. Full article (PDF). Reference page. .
Groenen, F. & A. Schreurs, 2012: Cenephasia (Cnephasiella) delnoyana Groenen & Schreurs, sp. n. from Spain (Lepidoptera: Tortricidae). Shilap Revista de Lepidopterologia 40 (159): 295–298.
Razowski, J. 1955. Eine neue südeuropäische Art der Gattung Cnephasia Curtis (Lepidoptera, Tortricidae). Zeitschrift Wiener Entomologische Gesellschaft 40: 264–266. Full article PDF. Reference page.
Razowski, J. 1958: New and little known Palaeartic species of the Cnepahsiini (Lepidoptera, Tortricidae). Acta zoologica cracoviensia 2 (25): 563-605. article.Reference page.
Razowski, J. 1959. Neue und wenig bekammte palaearktische Wickler-Arten (Lepidoptera, Tortricidae). Zeitschrift Wiener Entomologische Gesellschaft 44: 81-87. full article.Reference page.
Razowski, J. 1960. Remarks on the types of some African Tortricidae (Lepidoptera). Polish Journal of Entomologie 30(13): 169–172. Reference page.
Razowski, J. 1965. The Palaearctic Cnephasiini (Lepidoptera: Tortricidae). Acta zoologica cracoviensia 10(3): 199-343. full article (PDF). Reference page.
Razowski, J. 1983a: Descriptions of new Cnephasia Curtis (Tortricidae). Nota lepidopterologica (n.f.) 6(4): 235-238.Reference page.
Trematerra, P. 1991. Due nuovi Cnephasiini dai Monti del Pollino, Cnephasia zangheriana sp.n. e Cnephasia pollinoana sp.n. (Lepidoptera Tortricidae) Bolletino della Società entomologica italiana 123(2): 149–156. Reference page.
Zlatkov, B. 2013: Remarks on Tortricidae species with unknown and little known females. Nota Lepidopterologica 36(2): 191-197. Full article (PDF). Reference page.

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Superfamilia: Tortricoidea

Familia: Tortricidae
Subfamilia: Tortricinae
Tribus: Cnephasiini
Genus: Cnephasia
Subgenera: C. (Anoplocnephasia) – C. (Cnephasia) – C. (Cnephasiella) – C. (Hypostephanuncia)
Overview of assigned species

C. abrasana – C. daedalea – C. delnoyana – C. etnana – C. hellenica – C. heringi – C. incertana
Overview of species not assigned to a sub-genus

C. albatana – C. alfacarana – C. alticola – C. alticolana – C. amseli – C. asbolodes – C. asiatica – C. asseclana – C. bizensis – C. chrysantheana – C. cinereipalpana – C. communana – C. conspersana – C. constantinana – C. cupressivorana – C. disforma – C. disparana – C. divisana – C. ecullyana – C. ergastularis – C. facetana – C. finita – C. fiorii – C. fragosana – C. fulturata – C. galeotis – C. genitalana – C. graecana – C. grandis – C. gueneana – C. heinemanni – C. hunzorum – C. incinerata – C. josebai – C. jozefi – C. kasyi – C. kenneli – C. klimeschi – C. korvaci – C. laetana – C. lineata – C. longana – C. macrostoma – C. margelanensis – C. mediocris – C. microstrigana – C. minutula – C. nesiotica – C. nigripunctana – C. nigrofasciana – C. nowickii – C. olearis – C. orientana – C. oxyacanthana – C. palestinensis – C. parnassicola – C. pasiuana – C. personatana – C. phalarocosma – C. quentini – C. razowskii – C. regifica – C. sedana – C. semibrunneata – C. stachi – C. stephensiana – C. syriella – C. taganista – C. taurominana – C. tianshanica – C. tofina – C. tolli – C. tremewani – C. tripolitana – C. tristrami – C. ussurica – C. venusta – C. virginana – C. wimmeri – C. zangheriana – C. zelleri – C. zernyi
Overview of species not assigned to Cnephasia

C. catastrepta – C. chlorocrossa
Name

Cnephasia Curtis, 1826

Type species: Phalaena logiana Clerck
Synonyms

Anoplocnephasia Réal, 1953
Brachycnephasia Réal, 1953
Chephasia Razowski, 1977 (misspelling of Cnephasia)
Cnephasianella Benander, 1950
Cnephasiella Adamczewski, 1936
Cnephosia Desmarest, 1857 (misspelling of Cnephasia)
Hypostephanuncia Réal, 1951
Hypostephanuntia Razowski, 1977 (misspelling of Hypostephanuncia)
Sciaphila (Tortricidae) Treitschke, 1829

References

Alipanah, H. 2019. An overview of the tribe Cnephasiini (Lepidoptera: Tortricidae: Tortricinae) of Iran with description of a new species. Zootaxa 4661(3): 501–521. DOI: 10.11646/zootaxa.4661.3.5. Reference page.
Brown, J.W. 2005. World Catalogue of Insects. Volume 5: Tortricidae (Lepidoptera) 1–741. Apollo Books. Reference page.
Gastón, J., Vives Moreno, A. & Revilla, T. 2017. Descripción de tres especies nuevas de la Familia Tortricidae en la Península Ibérica (Lepidoptera: Tortricidae). SHILAP Revista de Lepidopterología 45(180): 689-698. Full article (PDF). Reference page. .
Groenen, F. & A. Schreurs, 2012: Cenephasia (Cnephasiella) delnoyana Groenen & Schreurs, sp. n. from Spain (Lepidoptera: Tortricidae). Shilap Revista de Lepidopterologia 40 (159): 295–298.
Razowski, J. 1955. Eine neue südeuropäische Art der Gattung Cnephasia Curtis (Lepidoptera, Tortricidae). Zeitschrift Wiener Entomologische Gesellschaft 40: 264–266. Full article PDF. Reference page.
Razowski, J. 1958: New and little known Palaeartic species of the Cnepahsiini (Lepidoptera, Tortricidae). Acta zoologica cracoviensia 2 (25): 563-605. article.Reference page.
Razowski, J. 1959. Neue und wenig bekammte palaearktische Wickler-Arten (Lepidoptera, Tortricidae). Zeitschrift Wiener Entomologische Gesellschaft 44: 81-87. full article.Reference page.
Razowski, J. 1960. Remarks on the types of some African Tortricidae (Lepidoptera). Polish Journal of Entomologie 30(13): 169–172. Reference page.
Razowski, J. 1965. The Palaearctic Cnephasiini (Lepidoptera: Tortricidae). Acta zoologica cracoviensia 10(3): 199-343. full article (PDF). Reference page.
Razowski, J. 1983a: Descriptions of new Cnephasia Curtis (Tortricidae). Nota lepidopterologica (n.f.) 6(4): 235-238.Reference page.
Trematerra, P. 1991. Due nuovi Cnephasiini dai Monti del Pollino, Cnephasia zangheriana sp.n. e Cnephasia pollinoana sp.n. (Lepidoptera Tortricidae) Bolletino della Società entomologica italiana 123(2): 149–156. Reference page.
Zlatkov, B. 2013: Remarks on Tortricidae species with unknown and little known females. Nota Lepidopterologica 36(2): 191-197. Full article (PDF). Reference page.


Cnephasia is a genus of tortrix moths (family Tortricidae). It belongs to the subfamily Tortricinae and therein to the tribe Cnephasiini, of which it is the type genus.[1]
Taxonomy and systematics

The type species around which Cnephasia was established by J. Curtis in 1826 (in the explanations to plate 100 of his British Entomology) was claimed to be "Tortrix logiana". However, this was a misidentification; the name of T. logiana (described by C.A. Clerck in 1759 and nowadays called Acleris logiana) was until the early 20th century frequently applied to related species in error.[2]

Curtis simply repeated the mistake of A.H. Haworth, who had in his 1811 volume of Lepidoptera Britannica discussed a Cnephasia under Clerck's name, but the original misidentification may well go back to Linnaeus' treatment of "T. logiana" in Systema naturae. Eventually, this was resolved, and the type species of Cnephasia was determined to be the tortrix moth described as Olethreutes pasiuana by J. Hübner in 1822.[3]
Species

The currently recognized species of Cnephasia are:[4]

Cnephasia abrasana (Duponchel in Godart, 1842)
Cnephasia albatana Chrétien, 1915
Cnephasia alfacarana Razowski, 1958
Cnephasia alticola Kuznetzov, 1966
Cnephasia alticolana
Cnephasia amseli (Lucas, 1942)
Cnephasia asiatica Kuznetzov, 1956
Cnephasia asseclana – flax tortrix
Cnephasia bizensis Réal, 1953
Cnephasia chrysantheana (Duponchel in Godart, 1842)
Cnephasia communana
Cnephasia conspersana Douglas, 1846
Cnephasia constantinana Razowski, 1958
Cnephasia cupressivorana (Staudinger, 1871)
Sciaphila debiliana Walker, 1863
Cnephasia delnoyana Groenen & Schreurs, 2012
Cnephasia disforma Razowski, 1983
Cnephasia disparana Kuznetzov in Danilevsky, Kuznetsov & Falkovitsh, 1962
Cnephasia divisana Razowski, 1959
Cnephasia ecullyana Réal, 1951
Cnephasia etnana Razowski & Trematerra, 1999
Cnephasia facetana Kennel, 1901
Cnephasia fiorii Razowski, 1958
Cnephasia fragosana (Zeller, 1847)
Cnephasia fulturata Rebel, 1940
Cnephasia genitalana Pierce & Metcalfe, 1915
Cnephasia graecana Rebel, 1902
Cnephasia grandis (Osthelder, 1938)
Cnephasia gueneana (Duponchel in Godart, 1836)
Cnephasia heinemanni Obraztsov, 1957
Cnephasia hellenica Obraztsov, 1956
Cnephasia heringi Razowski, 1958
Cnephasia hunzorum Diakonoff, 1971
Cnephasia incertana – light grey tortrix
Cnephasia jozefi Razowski, 1961
Cnephasia kasyi Razowski, 1971
Cnephasia kenneli Obraztsov, 1956

Cnephasia klimeschi Razowski, 1958
Cnephasia korvaci Razowski, 1965
Cnephasia laetana (Staudinger, 1871)
Cnephasia lineata (Walsingham, 1900)
Cnephasia longana (Haworth, [1811])
Cnephasia margelanensis Razowski, 1958
Cnephasia microstrigana Razowski, 1958
Cnephasia minutula Falkovitsh[verification needed] in Danilevsky, Kuznetsov & Falkovitsh, 1962
Cnephasia nesiotica Razowski, 1983
Cnephasia nigripunctana Amsel, 1959
Cnephasia nigrofasciana (Bruand, 1850)
Cnephasia nowickii Razowski, 1958
Cnephasia orientana (Alphéraky, 1877)
Cnephasia orthias Meyrick, 1910
Cnephasia oxyacanthana (Herrich-Schäffer, 1851)
Cnephasia parnassicola Razowski, 1958
Cnephasia pasiuana (Hübner, 1799)
Cnephasia personatana Kennel, 1901
Cnephasia razowskii Alipanah, 2019
Cnephasia regifica Razowski, 1971
Cnephasia sedana (Constant, 1884)
Cnephasia semibrunneata (de Joannis, 1891)
Cnephasia stachi Razowski, 1958
Cnephasia stephensiana – grey tortrix
Cnephasia tianshanica Filipjev, 1934
Cnephasia tofina Meyrick, 1922
Cnephasia tremewani Razowski, 1961
Cnephasia tripolitana Razowski, 1958
Cnephasia tristrami (Walsingham, 1900)
Cnephasia ussurica Filipjev, 1962
Cnephasia venusta Razowski, 1971
Cnephasia virginana (Kennel, 1899)
Cnephasia wimmeri Arenberger, 1998
Cnephasia zangheriana Trematerra, 1991
Cnephasia zelleri (Christoph, 1877)
Cnephasia zernyi Razowski, 1959

"Cnephasia" jactatana does not seem to belong in this genus.[4]
Former species

Cnephasia daedalea Razowski, 1983

Synonyms

Obsolete scientific names for Cnephasia are:[5]

Anoplocnephasia Réal, 1953
Brachycnephasia Réal, 1953
Chephasia (lapsus)
Cnephasianella (lapsus)
Cnephasiella Adamczewski, 1936
Cnephosia (lapsus)
Hypostephanuncia Réal, 1951
Hypostephanuntia (lapsus)
Sciaphila Treitschke in Ochsenheimer, 1829

Hypostephanuncia is sometimes listed as a junior synonym of the closely related genus Eana, but its type species is C. ecullyana. Whether the monotypic Sphaleroptera is a distinct and valid genus or merely a specialized offshoot of Cnephasia is not yet resolved; some authors include it here, while others don't.[6]
Footnotes

Baixeras et al. (2009a)
Curtis, [1826], Baixeras et al. (2009a), and see references in Savela (2010)
Curtis, [1826], Razowski (1998): 243
Baixeras et al. (2009b)
Baixeras et al. (2009a), and see references in Savela (2005a)

Baixeras et al. (2009a), and see references in Savela (2005a,b)

References

Alipanah, H. (2019): An overview of the tribe Cnephasiini (Lepidoptera: Tortricidae: Tortricinae) of Iran with description of a new species. Zootaxa 4661(3): 501-521.
Baixeras, J.; Brown, J.W. & Gilligan, T.M. (2009a): Online World Catalogue of the Tortricidae – Genus: Cnephasia account. Version 1.3.1. Retrieved 2009-JAN-20.
Baixeras, J.; Brown, J.W. & Gilligan, T.M. (2009b): Online World Catalogue of the Tortricidae – Cnephasia species list. Version 1.3.1. Retrieved 2009-JAN-20.
Curtis, John [1826]: [Genus: Cnephasia]. In: British Entomology, being illustrations and descriptions of the genera of insects found in Great Britain and Ireland, etc. 6(100): 1-2. E. Ellis & Co., London. Fulltext at the Internet Archive
Razowski, Józef (1998): The genera of Tortricidae. Part III. Nearctic Chlidanotinae and Tortricidae. Acta Zoologica Cracoviensia 41(2): 227-281. Fulltext at Google Books
Savela, Markku (2005a): Markku Savela's Lepidoptera and some other life forms – Cnephasia. Version of 2005-SEP-14. Retrieved 2010-APR-14.
Savela, Markku (2005b): Markku Savela's Lepidoptera and some other life forms – Eana. Version of 2005-SEP-14. Retrieved 2010-APR-14.
Savela, Markku (2010): Markku Savela's Lepidoptera and some other life forms – Acleris logiana. Version of 2010-FEB-19. Retrieved 2010-APR-14.

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