Cossyphodinae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Tenebrionoidea

Familia: Tenebrionidae
Subfamilia: Cossyphodinae
References

Latreille, P.A. 1802: Histoire naturelle, générale et particulière des Crustacés et des Insectes. Tome 3. F. Dufart, Paris. Internet Archive BHL
Schawaller, W. 2006: First record of the subfamily Cossyphodinae (Coleoptera: Tenebrionidae) from the Socotra Archipelago. Fauna of Arabia, 21: 247–250.
Schawaller, W. 2013: Cossyphodini (Coleoptera: Tenebrionidae: Pimeliinae) in South Africa, Namibia and adjacent regions: New species and records, key to genera, and Old World species catalogue. Zootaxa 3721(4): 351–364. DOI: 10.11646/zootaxa.3721.4.3 Reference page.