Cryptinae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Hymenopterida
Ordo: Hymenoptera
Subordo: Apocrita
Superfamilia: Ichneumonoidea

Familia: Ichneumonidae
Subfamilia: Cryptinae
Tribus: Claseini - Cryptini - Hemigasterini - Phygadeuontini
Overview of genera

Acrolyta – Afretha – Allophatnus – Amphibulus – Anepomias – Aptesis – Astomaspis – Ateleute – Atractodes – Bathythrix – Bodedia – Bozakites – Ceratocryptus – Ceratodolius – Ceratophenax – Ceratophygadeuon – Charitopes – Chirotica – Coccygodes – Costifrons – Cryptus – Cyanodolius – Cyanopelor – Diaglyptella – Diatora – Dichrogaster – Diracela – Dolichomastix – Encrateola – Eurycryptus – Fitatsia – Fotsiforia – Gabia – Gabunia – Gambroides – Gambrus – Gelis – Gerdius – Gessia – Goryphus – Gotra – Handaoia – Hemisphragia – Hoeocryptus – Indovia – Ischnus – Larpelites – Lavinifia – Lienella – Listrognathus – Madastenus – Mamelia – Mansa – Menaforia – Mesophragis – Necolio – Nematocryptus – Neopimpla – Nipponaetes – Odontocryptus – Owenus – Paraglyptus – Paraphylax – Perinetia – Phygadeuon – Piambia – Piasites – Picardiella – Platymystax – Pterocryptus – Rambites – Retalia – Rhynchocryptus – Savolia – Schreineria – Skeatia – Sozites – Spathacantha – Sphecoctonus – Stenarella – Synechocryptus – Tanyloncha – Tolonus – Trafana – Tricapus – Tsirambia – Xenolytus – Zonocryptus
Name

Cryptinae Kirby, 1837 [placed on the Official List of Family-Group Names in Zoology by ICZN (1994: 74)] [source: ICZN (1994: 74)]

Type genus: Cryptus Fabricius, 1804 [source: ICZN (1994: 74)] [stem = Crypt-]

References

ICZN 1994: OPINION 1757. Cryptus Fabricius, 1804 and CRYPTINAE Kirby, 1837 (Insecta, Hymenoptera): conserved. Bulletin of zoological nomenclature, 51: 74–75. ISSN: 0007-5167 Internet Archive BHL BioStor
Khalaim, A.I. 2008: Fossil ichneumon wasps (Hymenoptera: Ichneumonidae) from Biamo (Russia), Oligocene. Alavesia, 2: 101–112. PDF

Cryptinae is a subfamily of wasps in the family Ichneumonidae. The family has also been called Gelinae, Hemitelinae, and Phygadeuontinae by various authorities,[1] though the Phygadeuontinae have since been elevated to a separate subfamily.[2]

Description

Species are distributed worldwide. A 5-sided areolet, a short sternaulus, and a puffy face with a convex clypeus are diagnostic characters for most members of this large subfamily.[3]
Taxonomy

Gelinae was the name used by H. K. Townes in 1969.[4] Fitton and Gauld's review of ichneumonid families synonymized Gelinae under Hemitelinae in 1976[5] and later as Phygadeuontinae in 1978.[1] However, due to the decision of International Code of Zoological Nomenclature's Opinion 1715 in 1994 to conserve the status of the genus Cryptus Fabricius, 1804, both Gelinae and Phygadeuontinae became junior synonyms for Cryptinae.[6][2] Santos later restricted the family in 2017 to only include the tribes Aptesini and Cryptini, forming the separate families Claseinae and Phygadeuontinae for the remaining tribes of Claseini (1 genus) and Phygadeuontini (123 genera), respectively.[2]

There are about 273 genera in Cryptinae between the tribes of Aptesini and Cryptini.[7][8]
Classification

The following classification of Phygadeuontini is based on Wahl (2014):[8]
Tribe Cryptini
Agrothereutes abbreviatus, female

Subtribe Agrothereutina
Agrothereutes Förster, 1850 — Holarctic
Amauromorpha Ashmead, 1905 — Oriental
Apsilops Förster, 1869 — Holarctic
Aritranis Förster, 1869 — Holarctic
Gambrus Förster, 1869 — Holarctic
Hidryta Förster, 1869 — Holarctic
Idiolispa Förster, 1869 — Holarctic
Mesostenidea Viereck, 1914 — Palearctic
Pycnocryptodes Aubert, 1971 — Palearctic
Pycnocryptus Thomson, 1873 — Holarctic
Thrybius Townes, 1965 — Palearctic
Trychosis Förster, 1869 — Holarctic, Oriental
Subtribe Ateleutina
Ateleute Förster, 1869 — worldwide
Tamaulipeca Kasparyan & Hernandez, 2001 — Neotropical
Subtribe Baryceratina
Baryceros Gravenhorst, 1829 — Nearctic, Neotropical
Brussinocryptus Pagliano & Scarramozino, 1990 — Oriental, Palearctic
Buysmania Cheesman, 1941 — Oriental
Calaminus Townes, 1965 — Palearctic
Ceratomansa Cushman, 1922 — Australian
Chlorocryptus Cameron, 1903 — Oriental, Palearctic
Coccygodes Saussure, 1892 — Ethiopian
Jonathania Gupta, 1987 — Oriental
Lamprocryptidea Viereck, 1913 — Neotropical
Paragambrus Uchida, 1936 — Palearctic
Prionacis Townes, 1970a — Oriental
Whymperia Cameron, 1903 — Nearctic, Neotropical
Subtribe Ceratocryptina
Acromia Townes, 1961b — Oriental
Afretha Seyrig, 1952 — Ethiopian
Aprix Townes, 1961b — Australian
Ceratocryptus Cameron, 1903 — Ethiopian, Oriental
Ceratodolius Seyrig, 1952 — Ethiopian
Chamula Townes, 1962 — Nearctic, Neotropical
Cheesmanella Ozdikmen, 2005 — Oriental
Cremnocryptus Cushman, 1945 — Australian, Oriental
Ecaepomia Wang, 2001 — Oriental
Fitatsia Seyrig, 1952 — Ethiopian, Oriental
Lipoprion Townes, 1970a — Oriental
Lorio Cheesman, 1936 — Oriental
Malaycromia Gupta, 1999 — Oriental
Neaprix Gauld, 1984 — Australian
Nematocryptus Roman, 1910 — Ethiopian, Oriental
Piasites Seyrig, 1952 — Ethiopian
Rambites Seyrig, 1952 — Ethiopian
Silsila Cameron, 1903 — Oriental
Thelodon Townes, 1961b — Australian, Oriental
Trachyglutus Townes, 1970a — Neotropical
Wuda Cheesman, 1936 — Oriental
Subtribe Coesulina
Coesula Cameron, 1905 — Oriental
Subtribe Cryptina

Cryptus albitarsis, female

Aeglocryptus Porter, 1987 — Neotropical
Aeliopotes Porter, 1985 — Neotropical
Aglaodina Porter, 1987 — Neotropical
Anacis Porter, 1967 — Australian, Neotropical
Araucacis Porter, 1987 — Neotropical
Biconus Townes, 1970a — Neotropical
Buathra Cameron, 1903 — Holarctic, Oriental
Caenocryptoides Uchida, 1936 — Oriental, Palearctic
Caenocryptus Thomson, 1873 — Holarctic
Caenopelte Porter, 1967 — Neotropical
Camera Townes, 1962 — Nearctic, Neotropical
Chilecryptus Porter, 1987 — Neotropical
Chromocryptus Ashmead, 1900 — Neotropical
Compsocryptus Ashmead, 1900 — Nearctic, Neotropical
Cosmiocryptus Cameron, 1902 — Neotropical (Porter, 1987)
Cryptopteryx Ashmead, 1900 — Neotropical
Cryptus Fabricius, 1804 — Ethiopian, Holarctic, Oriental
Cyanodolius Seyrig, 1952 — Ethiopian
Cyanopelor Townes, 1973 — Ethiopian
Cyclaulus Townes, 1970a — Neotropical
Dihelus Townes, 1970a — Oriental, Palearctic
Diplohimas Townes, 1970a — Neotropical
Distictus Townes, 1966 — Neotropical
Dochmidium Porter, 1967 — Neotropical (Porter, 1987)
Dotocryptus Brèthes, (1918) 1919 — Neotropical
Enclisis Townes, 1970a — Palearctic
Etha Cameron, 1903 — Oriental
Gessia Townes, 1973 — Ethiopian
Glabridorsum Townes, 1970a — Australian, Oriental, Palearctic
Gyropyga Townes, 1970a — Oriental, Palearctic
Hedycryptus Cameron, 1903 — Oriental, Palearctic
Hypsanacis Porter, 1987 — Neotropical
Ischnus Gravenhorst, 1829 — worldwide
Itamuton Porter, 1987 — Neotropical
Joppidium Cresson, 1872 — Nearctic, Neotropical
Lanugo Townes, 1962 — Nearctic, Neotropical
Leptarthron Townes, 1970a — Neotropical
Meringopus Förster, 1869 — Holarctic
Mesophragis Seyrig, 1952 — Ethiopian
Monothela Townes, 1970a — Neotropical
Myrmecacis Porter, 1987 — Neotropical
Myrmeleonostenus Uchida, 1936 — Australian, Oriental, Palearctic
Nebostenus Gauld, 1984 — Australian
Nelophia Porter, 1967 — Neotropical
Neocryptopteryx Blanchard, 1947 — Neotropical (Porter, 1987)
Neodontocryptus Uchida, 1940 — Oriental
Nippocryptus Uchida, 1936 — Oriental, Palearctic
Nothischnus Porter, 1967 — Neotropical (Porter, 1987)
Odontocryptus Saussure, 1892 — Ethiopian
Oecetiplex Porter, 1987 — Neotropica
Palmerella Cameron, 1908 — Oriental, Palearctic
Periplasma Porter, 1967 — Neotropical (Porter, 1987)
Phycitiplex Porter, 1987 — Neotropical
Picrocryptoides Porter, 1965 — Neotropical
Reptatrix Townes, 1962 — Nearctic
Rhynchocryptus Cameron, 1905 — Ethiopian (Townes & Townes, 1973)
Sciocryptus Porter, 1987 — Neotropical
Synechocryptus Schmiedeknecht, 1904 — Ethiopian, Palearctic
Trachysphyrus Haliday, 1836 — Neotropical
Tricentrum Townes, 1970a — Neotropical
Trihapsis Townes, 1970a — Neotropical
Xiphonychidion Porter, 1967 — Neotropical (Porter, 1987)
Xylacis Porter, 1987 — Neotropical
Xylophrurus Förster, 1869 — Holarctic
Xylostenus Gauld, 1984 — Australian
Zonocryptus Ashmead, 1900 — Ethiopian
Subtribe Gabuniina

Xanthocryptus novozealandicus, female

Agonocryptus Cushman, 1929 — Nearctic, Neotropical
Ahilya Gupta & Gupta, 1985 — Oriental
Amrapalia Gupta & Jonathan, 1970 — Oriental
Anepomias Seyrig, 1952 — Ethiopian
Apocryptus Uchida, 1932 — Oriental
Arhytis Townes, 1970a — Australian, Oriental
Cestrus Townes, 1966 — Neotropical
Cryptohelcostizus Cushman, 1919 — Nearctic
Dagathia Cameron, 1903 — Oriental
Digonocryptus Viereck, 1913 — Neotropical
Eurycryptus Cameron, 1901 — Australian, Ethiopian, Oriental
Fenixia Aguiar, 2005 — Neotropical
Fortipalpa Kasparyan & Ruiz, 2007 — Neotropical
Gabunia Kriechbaumer, 1895 — Ethiopian
Gerdius Townes, 1970a — Ethiopian
Hackerocryptus Gauld, 1984 — Australian
Hadrocryptus Cameron, 1903 — Oriental
Kemalia Koçak, 2009 — Oriental
Kriegeria Ashmead, 1905 — Oriental (Gupta, 1987)
Lagarosoma Gupta & Gupta, 1984 — Neotropical
Lophoglutus Gauld, 1984 — Australian
Microstenus Szépligeti, 1916 — Oriental
Nesolinoceras Ashmead, 1906 — Neotropical
Pharzites Cameron, 1905 — Oriental
Prosthoporus Porter, 1976 — Neotropical
Pterocryptus Szépligeti, 1916 — Ethiopian, Palearctic
Schreineria Schreiner, 1905 — Ethiopian, Oriental, Palearctic
Spathacantha Townes, 1970a — Ethiopian
Tanepomidos Gupta & Jonathan, 1971 — Oriental
Torbda Cameron, 1902 — Oriental
Trypha Townes, 1970a — Neotropical
Xanthocryptus Cameron, 1901 — Australian, Oriental
Xoridesopus Cameron, 1907 — Oriental
Subtribe Glodianina
Dicamixus Szépligeti, 1916 — Neotropical
Glodianus Cameron, 1902 — Neotropical
Lamprocryptus Schmiedeknecht, 1904 — Neotropical
Subtribe Goryphina
Allophatnus Cameron, 1905 — Australian, Ethiopian, Oriental (Townes, 1971)
Baltazaria Townes, 1961b — Neotropical, Oriental
Bozakites Seyrig, 1952 — Ethiopian
Buodias Cameron, 1902 — Ethiopian, Oriental (Gupta, 1987)
Hoeocryptus Habermehl, 1902
Takastenus Uchida, 1931 (Gauld, 1984 — Gupta, 1987)
Soratsia Seyrig, 1952 (Townes & Townes, 1973)
Ceratella Seyrig, 1952
Calosphyrum Townes, 1970a — Oriental
Ceratophenax Seyrig, 1952 — Ethiopian
Colaulus Townes, 1970a — Oriental
Costifrons Townes, 1970a — Ethiopian
Debilos Townes, 1966 — Neotropical
Diapetimorpha Viereck, 1913 — Nearctic, Neotropical
Euchalinus Townes, 1961b — Australian, Oriental
Formostenus Uchida, 1931 (Jonathan, 1980) — Oriental
Fotsiforia Seyrig, 1952 — Ethiopian, Oriental (Jonathan, 1980)
Friona Cameron, 1902 — Oriental
Gambroides Betrem, 1941 — Australian, Ethiopian, Oriental (Jonathan, 1980)
Goryphus Holmgren, 1868 — Australian, Ethiopian, Oriental
Hemisphragia Seyrig, 1952 — Ethiopian
Hoeocryptus Habermehl, 1902 — Ethiopian
Hylophasma Townes, 1970a — Holarctic, Neotropical
Isotima Förster, 1869 — Oriental, Ethiopian
Larpelites Cameron, 1904 — Ethiopian
Lavinifia Seyrig, 1952 — Ethiopian
Listrognathus Tschek, 1870 — Australian, Ethiopian, Holarctic, Oriental
Loxopus Townes, 1970a — Neotropical
Madagascesa Kocak & Kemal, 2008 - formerly Perinetia Seyrig, 1952 — Ethiopian
Madastenus Seyrig, 1952 — Ethiopian
Melcha Cameron, 1902 — Oriental
Menaforia Seyrig, 1952 — Ethiopian, Oriental
Necolio Cheesman, 1936 — Australian, Ethiopian, Oriental, Palearctic
Neobuodias Rao & Nikam, 1984 — Oriental
Owenus Townes, 1970a — Ethiopian
Perjiva Jonathan & Gupta, 1973 — Oriental
Piambia Seyrig, 1952 — Ethiopian
Pseudotricapus Jonathan, 1987 — Oriental
Skeatia Cameron, 1901 — Ethiopian, Oriental (Jonathan & Gupta, 1973)
Syntrips Gauld, 1984 — Australian
Tanyloncha Townes, 1970a — Ethiopian
Tolonus Seyrig, 1952 — Ethiopian
Trafana Seyrig, 1952 — Ethiopian
Tricapus Townes, 1970a — Ethiopian
Tsirambia Seyrig, 1952 — Ethiopian
Subtribe Lymeonina

Lymeon orbus, female

Acerastes Cushman, 1929 — Nearctic, Neotropical
Basileucus Townes, 1970a — Neotropical
Bathyzonus Townes, 1970a — Neotropical
Bicryptella Strand, 1917 — Neotropical
Dismodix Townes, 1966 — Neotropical
Golbachiella Townes, 1970a — Neotropical
Latosculum Townes, 1966 — Neotropical
Lymeon Förster, 1869 — Nearctic, Neotropical
Mallochia Viereck, 1912 — Nearctic, Neotropical
Pachysomoides Strand, 1917 — Nearctic, Neotropical
Petila Tedesco & Aguiar, 2009 — Neotropic
Polycyrtidea Viereck, 1913 — Nearctic, Neotropical
Polyphrix Townes, 1970a — Neotropical
Priotomis Townes, 1970a — Neotropical
Rhinium Townes, 1966 — Neotropical
Savolia Seyrig, 1952 — Ethiopian
Strabotes Townes, 1970a — Neotropical
Toechorychus Townes, 1946 — Neotropical
Subtribe Melanocryptina
Melanocryptus Cameron, 1902 — Neotropical
Subtribe Mesostenina

Mesostenus thoracicus, female

Acorystus Townes, 1970a — Neotropical
Anupama Jonathan, 1982 — Oriental
Bicristella Townes, 1966 — Neotropical
Cryptanura Brullé, 1846 — Nearctic, Neotropical
Diloa Cheesman, 1936 — Australian, Oriental
Gotra Cameron, 1902 — Ethiopian, Oriental
Gyrolaba Townes, 1970a — Oriental
Harpura Townes, 1970a — Neotropical
Hercana Townes, 1970a — Neotropical
Irabatha Cameron, 1906 — Australian, Oriental
Junctivena Gauld, 1984 — Australian
Mecistum Townes, 1970a — Neotropical
Mesostenus Gravenhorst, 1829 — worldwide
Paranacis Gauld, 1984 — Australian
Polycyrtus Spinola, 1840 — Nearctic, Neotropical
Stiromesostenuss Cameron, 1911 — Australian, Oriental
Tomagotras Gauld, 1984 — Australian
Tretobasis Porter, 1973 — Neotropical
Subtribe Osprynchotina
Acroricnus Ratzeburg, 1852 — Holarctic
Iaria Cheesman, 1936 — Australian, Oriental
Messataporus Cushman, 1929 — Nearctic, Neotropical
Nematopodius Gravenhorst, 1829 — Oriental, Palearctic
Osprynchotus Spinola, 1841 — Ethiopian
Photocryptus Viereck, 1913 — Neotropical
Poecilopimpla Morley, 1914;
Picardiella Lichtenstein, 1920 — Ethiopian, Oriental, Palearctic
Sphecoctonus Seyrig, 1952 — Ethiopian
Stenarella Szépligeti, 1916 — Australian, Ethiopian, Oriental
Subtribe Sphecophagina
Arthula Cameron, 1900 — Australian, Oriental, Palearctic
Latibulus Gistel, 1848 — Oriental, Palearctic
Sphecophaga Westwood, 1840 — Holarctic
Subtribe Vagenathina
Stetholophus Townes, 1970a — Oriental
Vagenatha Cameron, 1901 — Oriental

Tribe Aptesini
Aptesis nigrocincta, female

Aconias Cameron, 1904 — Palearctic, Oriental
Aptesis Förster, 1850 — Ethiopian, Holarctic, Oriental
Colocnema Förster, 1869 — Palearctic
Cratocryptus Thomson, 1873 — Holarctic
Cubocephalus Ratzeburg, 1848 — Holarctic
Demopheles Förster, 1869 — Holarctic, Oriental
Echthrus Gravenhorst, 1829 — Holarctic
Giraudia Förster, 1869 — Holarctic
Hemigaster Brullé, 1846 — Oriental
Javra Cameron, 1903 — Holarctic, Oriental
Listrocryptus Brauns, 1905 — Palearctic
Litochila Momoi, 1965 — Oriental, Palearctic
Livipurpurata Wang & Yao, 1994 — Oriental
Mansa Tosquinet, 1896 — Ethiopian, Oriental
Megaplectes Förster, 1869 — Holarctic
Notocampsis Townes, 1970a — Neotropical
Oresbius Marshall, 1867 — Holarctic
Oxytaenia Förster, 1869 — Nearctic, Neotropical
Parmortha Townes, 1962 — Holarctic
Platymystax Townes, 1970a — Ethiopian, Neotropical, Oriental
Plectrocryptus Thomson, 1874 — Oriental, Palearctic
Pleolophus Townes, 1962 — Holarctic
Polytribax Förster, 1869 — Holarctic, Neotropical, Oriental
Rhytura Townes, 1962 — Nearctic
Schenkia Förster, 1869 — Holarctic
Stomacis Townes, 1970a — Oriental

References

Fitton, M.G; Gauld, I.D. (1978). "Further notes on family-group names of the Ichneumonidae (excluding Ichneumoninae) (Hymenoptera)". Systematic Entomology. 3 (3): 245–247. doi:10.1111/j.1365-3113.1978.tb00118.x. S2CID 84104340. Retrieved December 29, 2021.
Santos, B.F. (2017). "Phylogeny and reclassification of Cryptini (Hymenoptera, Ichneumonidae, Cryptinae), with implications for ichneumonid higher-level classification". Systematic Entomology. 42 (4): 650–676. doi:10.1111/syen.12238. S2CID 90996975. Retrieved December 29, 2021.
Cryptinae. WaspWeb. Iziko Museums.
Townes, Henry K. (1969). "Genera of Ichneumonidae, Part 2 (Gelinae)". Memoirs of the American Entomological Institute. 12: 1–537.
Fitton, M.G; Gauld, I.D. (1976). "The family-group names of the Ichneumonidae (excluding Ichneumoninae) (Hymenoptera)". Systematic Entomology. 1: 247–258.
"Opinion 1757. Cryptus Fabricius, 1804 and Cryptinae Kirby, 1837 (Insecta, Hymenoptera): conserved". Bulletin of Zoological Nomenclature. IZCN. 51: 74–75. 1994. ISSN 0007-5167. Retrieved December 29, 2021.
Broad, G., & Barthélémy, C. (2012). A new species of Hadrocryptus (Hymenoptera: Ichneumonidae: Cryptinae), with the first account of the biology for the genus. Journal of Hymenoptera Research, 24, 47.
Wahl, David (2014). "Synoptic Lists of World Genera and Family-group Names". American Entomological Institute. Retrieved December 29, 2021.