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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Blattodea

Familia: Ectobiidae
Subfamilia: Ectobiinae
Genus: Ectobius
Species: E. aetnaeus – E. aeoliensis – E. brunneri – E. eckerleini – E. filicensis – E. lagrecai – E. lapponicus – E. lucidus – E. pallidus – E. parvosacculatus – E. pyrenaicus – E. siculus – E. sylvestris – E. usticaensis – †E. kohlsi – ...
Name

Ectobius Stephens, 1835

References

Bohn, H. 1989: Revision of the sylvestris group of Ectobius Stephens in Europe (Blattaria: Blattellidae). Entomologica scandinavica, 20: 317–342. DOI: 10.1163/187631289X00357


Ectobius is a genus of non-cosmopolitan cockroaches once thought native to the Old World and described by Stephens in 1835,[1] belonging to the family Ectobiidae, subfamily Ectobiinae.[2] The discovery[3] of 4 ectobius cockroaches in Colorado dating to 49 Million years ago suggests the genus actually originated in North America. This genus has been subject to a number of revisions.[4][5]

Description
Ectobius species – Nymph

The adult 'cockroaches' reach 6–12 millimetres (0.24–0.47 in) of length, the basic coloration of their body is mostly brown or yellowish, with a clearer margin.

The females are usually bigger than the males and have shorter wings, while in the males wings cover at least the whole abdomen.
Distribution

Species of this genus are mainly present in most of Europe, Africa, in eastern Palearctic realm and the Near East. The "lesser cockroach" (i.e. as found in Britain), previously placed here, is now in the genus Capraiellus.[5]

The genus was long absent from the North American continent until recent re-introductions of some cool-adapted species in the Canadian Maritimes and North Eastern, US.[6]
Species
Ectobius lapponicus
Ectobius pallidus
Ectobius sylvestris
Ectobius vittiventris

The Cockroach Species File lists:[2]

Ectobius hipposiderus Bohn, 2013

subgenus Ectobiola Uvarov, 1940 (synonym Ectobiella Adelung, 1917)

Ectobius duskei Adelung, 1906

subgenus Ectobius Stephens, 1835

Ectobius aeoliensis Failla & Messina, 1974
Ectobius aethiopicus (Shelford, 1910)
Ectobius aetnaeus Ramme, 1927
Ectobius africanus Saussure, 1899
Ectobius albicinctus (Brunner von Wattenwyl, 1861)
Ectobius alleni Rehn, 1931
Ectobius baccettii Failla & Messina, 1979
Ectobius balcani Ramme, 1923
Ectobius brunneri Seoane, 1879
Ectobius burri Adelung, 1917
Ectobius corsorum Ramme, 1923
Ectobius darbandae Rehn, 1931
Ectobius delicatulus Bey-Bienko, 1950
Ectobius eckerleini Harz, 1977
Ectobius erythronotus Burr, 1898
Ectobius filicensis Failla & Messina, 1974
Ectobius frieseanus Princis, 1963
Ectobius heteropterus Bey-Bienko, 1963
Ectobius ichnusae Failla & Messina, 1982
Ectobius indicus Bey-Bienko, 1938
Ectobius intermedius Failla & Messina, 1981
Ectobius involutus Rehn, 1931
Ectobius jarringi (Hanitsch, 1937)
Ectobius kervillei Bolívar, 1907
Ectobius kikensis Rehn, 1931
Ectobius kikuyuensis Rehn, 1931
Ectobius kirgizius Bey-Bienko, 1936
Ectobius kraussianus Ramme, 1923
Ectobius lagrecai Failla & Messina, 1981
Ectobius lapponicus (Linnaeus, 1758) - type species (as Blatta lapponica L. = Ectobius lapponicus lapponicus)
Ectobius larus Rehn, 1931
Ectobius leptus Rehn, 1931
Ectobius lineolatus (Rehn, 1922)
Ectobius lodosi Harz, 1983
Ectobius lucidus (Hagenbach, 1822)
Ectobius makalaka Rehn, 1931
Ectobius minutus Failla & Messina, 1978
Ectobius montanus Costa, 1866
Ectobius neavei Shelford, 1911
Ectobius nuba Rehn, 1931
Ectobius pallidus (Olivier, 1789)
Ectobius palpalis Chopard, 1958
Ectobius parvosacculatus Failla & Messina, 1974
Ectobius pavlovskii Bey-Bienko, 1936
Ectobius punctatissimus Ramme, 1922
Ectobius pusillus Bey-Bienko, 1967
Ectobius pyrenaicus Bohn, 1989
Ectobius rammei Rehn, 1926
Ectobius sardous Baccetti, 1991
Ectobius scabriculus Failla & Messina, 1976
Ectobius semenovi Bey-Bienko, 1935
Ectobius siculus Ramme, 1949
Ectobius sjoestedti (Shelford, 1910)
Ectobius stanleyanus Rehn, 1931
Ectobius subvitreus Bey-Bienko, 1936
Ectobius supramontes Bohn, 2004
Ectobius sylvestris (Poda, 1761)
Ectobius tadzhikus Bey-Bienko, 1935
Ectobius textilis Rehn, 1931
Ectobius ticinus Bohn, 2004
Ectobius togoensis Ramme, 1923
Ectobius tuscus Galvagni, 1978
Ectobius tyrrhenicus Failla & Messina, 1973
Ectobius usticaensis Failla & Messina, 1974
Ectobius vittiventris (Costa, 1847)
Ectobius willemsei Failla & Messina, 1980

References

Stephens (1835) Illustrations of British Entomology. Mandibulata. Baldwin & Cradock, London, vol. VII, p. 1–306.
Cockroach Species File (retrieved 6 January 2021)
"After a 49-million-year hiatus, a cockroach reappears in North America". phys.org. Retrieved 2021-06-28.
Bohn, H. 1989: Revision of the sylvestris group of Ectobius Stephens in Europe (Blattaria: Blattellidae). Entomologica Scandinavica, 20: 317–342.
Bohn (2013) In Bohn, Beccaloni, Dorow & Pfeifer. Another species of European Ectobiinae travelling north – the new genus Planuncus and its relatives (Insecta: Blattodea: Ectobiinae). Arthropod Systematics & Phylogeny 71(3): 147.

Vršanský, P.; Oružinský, R.; Barna, P.; Vidlička, L'.; Labandeira, C. C. (2014). "Native Ectobius (Blattaria: Ectobiidae) From the Early Eocene Green River Formation of Colorado and Its Reintroduction to North America 49 Million Years Later". Annals of the Entomological Society of America. 107 (1): 28–36. doi:10.1603/AN13042. ISSN 0013-8746.

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