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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Subfamilia: Euschemoninae
Genus: Euschemon

Euschemoninae Kirby, 1897.

Euschemonidae Kirby, 1897: 33.
Euschemoninae (Kirby); Warren, Ogawa, Brower, 2009.


Kirby, W.F., 1896-97. A hand-book to the order Lepidoptera. Edward Lloyd, London. Vol.1, Vol.2, Vol.3, Vol.4, Vol.5. Reference page.
Li, W., Cong. Q., Shen, J.H., Zhang, J., Hallwachs, W., Janzen, D.H. & Grishin, N.V. 2019. Genomes of skipper butterflies reveal extensive convergence of wing patterns. PNAS 116(13): 6232–6237. DOI: 10.1073/pnas.1821304116. Text. SI Appendix. Reference page.
Sahoo, R.K., Warren, A.D., Wahlberg, N., Brower, A.V.Z., Lukhtanov, V.A. & Kodandaramaiah, U. 2016. Ten genes and two topologies: an exploration of higher relationships in skipper butterflies (Hesperiidae). PeerJ 4: e2653. DOI: 10.7717/peerj.2653 Reference page.
Toussaint, E.F.A., Breinholt, J.W., Earl, C., Warren, A.D., Brower, A.V.Z., Yago, M., Dexter, K.M., Espeland, M., Pierce, N.E., Lohman, D.J. & Kawahara, A.Y. 2018. Anchored phylogenomics illuminates the skipper butterfly tree of life. BMC Evolutionary Biology 18: 101. DOI: 10.1186/s12862-018-1216-z Reference page.
Warren, A.D., Ogawa, J.R. & Brower, A.V.Z. 2009. Revised classification of the family Hesperiidae (Lepidoptera: Hesperioidea) based on combined molecular and morphological data. Systematic entomology 34(3): 467–523. Reference page.
Zhang, J., Cong. Q., Shen, J.H., Brockmann, E. & Grishin, N.V. 2019. Three new subfamilies of skipper butterflies (Lepidoptera, Hesperiidae). ZooKeys 861: 91–105. DOI: 10.3897/zookeys.861.34686 Reference page.

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