Graphoderus

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Adephaga

Familia: Dytiscidae
Subfamilia: Dytiscinae
Tribus: Aciliini
Genus: Graphoderus
Species: G. adamsii – G. austriacus – G. bieneri – G. bilineatus – G. cinereus – G. perplexus – G. verrucifer – G. zonatus
Name

Graphoderus Dejean, 1833: 54 (as "Graphoderus. Eschscholtz.")

Type species: Dytiscus cinereus Linnaeus, 1758, by subsequent designation by Westwood, 1838: 8.
References
Primary references

Dejean, P.F.M.A. 1833. Catalogue des Coléoptères de la collection de M. le Comte Dejean. Deuxième édition. [Livraison 1 (pp. 1–96), Livraison 2 (pp. 97–176)]. Paris: Méquignon-Marvis Père et Fils. [pp. 177–256, Livraison 3 was published in 1834; pp. 257–360, Livraison 4 in 1835; pp. 361–442, Livraison 5 in 1836]. BHL. Reference page.

Additional references

Holmgren, S., Angus, R.B., Jia, F-L., Chen, Z-N. & Bergsten, J. 2016. Resolving the taxonomic conundrum in Graphoderus of the east Palearctic with a key to all species (Coleoptera, Dytiscidae). ZooKeys 574: 113–142. DOI: 10.3897/zookeys.574.7002.Reference page.

Links

ION
Nomenclator Zoologicus

Vernacular names
русский: Подводень

Graphoderus is a genus of beetle in family Dytiscidae native to the Holarctic.[1]

Extant species

The genus Graphoderus contains the following 12 species:[2]

Graphoderus adamsii (Clark, 1864)
Graphoderus austriacus (Sturm, 1834)
Graphoderus bieneri Zimmermann, 1921
Graphoderus bilineatus (De Geer, 1774)
Graphoderus cinereus (Linnaeus, 1758)
Graphoderus elatus Sharp, 1882
Graphoderus fascicollis (Harris, 1828)
Graphoderus liberus (Say, 1825)
Graphoderus manitobensis Wallis, 1933
Graphoderus occidentalis Horn, 1883
Graphoderus perplexus Sharp, 1882
Graphoderus zonatus (Hoppe, 1795)

Extinct species

These two extinct species are known only from fossils:[2]

†Graphoderus heeri Nilsson, 2001
†Graphoderus mirabilis Riha, 1974

References

Miller, K.B.; J. Bergsten (2016). Diving Beetles of the World: Systematics and Biology of the Dytiscidae. pp. 131–132. ISBN 978-1-4214-2054-7.
Nilsson, Anders N.; Hájek, Jiří (2021). "A World Catalogue of the Family Dytiscidae or the Diving Beetles (Coleoptera, Adephaga)" (PDF) (1.I.2021 ed.).