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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Lycaenidae
Subfamilia: Theclinae
Tribus: Iolaini
Genus: Iolaus
Subgenera (9):
I. (Aphniolaus) – I. (Argiolaus) – I. (Epamera) – I. (Iolaphilus) – I. (Iolaus) – I. (Philiolaus) – I. (Pseudiolaus) – I. (Tanuetheira) – I. (Trichiolaus)
Overview of species

per Stempffer, 1967 & Williams, 2007.
I. adorabilis – I. aemulus – I. aethria – I. aequatorialis – I. agnes – I. alcibiades – I. alienus – I. anesius – I. apatosa – I. aphnaeoides – I. arborifera – I. argentarius – I. aurivillii – I. australis – I. bakeri – I. banco – I. bansana – I. belli – I. bellina – I. bergeri – I. bilineata – I. bolissus – I. caesareus – I. calisto – I. carina – I. christofferi – I. coelestis – I. cottrelli – I. crawshayi – I. creta – I. cytaeis – I. diametra – I. dianae – I. djaloni – I. dubiosa – I. eurisus – I. farquharsoni – I. flavilinea – I. fontainei – I. frater – I. gabunica – I. gemmarius – I. glaucus – I. handmani – I. helenae – I. hemicyanus – I. henryi – I. iasis – I. icipe – I. ismenias – I. iturensis – I. iulus – I. jacksoni – I. jadwigae – I. jamesoni – I. kayonza – I. kelle – I. kupe – I. laon – I. laonides – I. lalos – I. liberiana – I. likpe – I. longicauda – I. lukabas – I. maesa – I. manasei – I. mane – I. maritimus – I. menas – I. mermeros – I. mermis – I. mimosae – I. montana – I. moyambina – I. ndolae – I. neavei – I. newporti – I. nolaensis – I. nursei – I. obscurus – I. ofere – I. pallene – I. pamae – I. parasilanus – I. paneperata – I. penningtoni – I. piaggiae – I. poecilaon – I. pollux – I. poultoni – I. pseudofrater – I. pseudopollux – I. sappirus – I. schultzei – I. scintillans – I. sciophilus – I. shaba – I. sibella – I. sidus – I. silanus – I. silarus – I. silas – I. stenogrammica – I. stewarti – I. sudanicus – I. tajoraca – I. theodori – I. timon – I. trimeni – I. timon – I. vansomereni – I. violacea

Name

Iolaus Hübner, [1819]: 81.

Type species: Papilio eurisus Cramer, [1780]
Synonymy

Jolaus Auctt. Mis-spelling.
Epamera Druce, 1891: 139.
Argiolaus Druce, 1891: 139.
Tanuetheira Druce, 1891: 139.
Jalous [sic] Hübner; Pagenstecher, 1897: 419, [mis-spelling?].
Trichiolaus Aurivillius, 1898: 317.
Aphniolaus Druce, 1902 [1903]: 117.
Pseudiolaus Riley, 1928: 392.
Iolaphilus Stempffer & Bennett, 1958: 1298.
Philiolaus Stempffer & Bennett, 1958: 1336.

References

Aurivillius, C. [1899]. Rhopalocera æthiopica; die Tagfalter des æthiopischen Faunengebietes… Bihang till Kungliga Svenska vetenskapsakademiens handlingar 31(5): 1–561. BHL. DOI: 10.5962/bhl.title.8773 Reference page.
Bayliss, J.L., Collins, S.C. & Congdon, T.C.E. 2016. A new species of Iolaus Hübner, [1819] subgenus Epamera Druce, 1891 (Lepidoptera: Lycaenidae: Theclinae) from Mts Namuli and Mabu, Northern Mozambique. Metamorphosis. 27: 23–30. Full article (PDF). Reference page.
Collins, S. C. & Larsen, T.B. 1995b. A new Iolaus from the Shaba province of Zaire (Lepidoptera; Lycaenidae). Lambillionea 95(3): 563–564. full article (PDF). Reference page.
Druce, H.H. 1891. On some African Butterflies hitherto referred to the Genus Iolaus, with Descriptions of new Species. Annals and Magazine of Natural History (6) 8: 139–150. BHL. Reference page.
Druce, H.H. 1902. On some new and little-known butterflies of the family Lycaenidae from the African, Australian and Oriental Regions. Proceedings of the Zoological Society of London 1902(2): 112–121, pls. 11,12. text, plates. Reference page.
Heath, A., 1985. Revision of three Iolaus subgenera and descriptions of two new species of Lycaenidae (Lepidoptera) from Zambia. Arnoldia Zimbabwe 9(18): 257–266. Reference page.
Hübner, J. 1819. Verzeichniß bekannter Schmettlinge 2–11: 17–176. BHL Reference page.
Larsen, T.B., 2005. Butterflies of West Africa. 2 vols, Apollo Books, Svendborg, Denmark: 1–595, 1-270. Reference page.
Pagenstecher, A. F., 1897. In Kükenthal, W. Ergebnisse einer zoologischen Forschungsreise in den Molukken und in Borneo. Lepidopteren. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 23: 353-467, pls 18-20. Reference page.
Riley, N.D., 1928. Notes on Iolaus, Argiolaus and related genera with descriptions of new species, subspecies and a new genus. (Lepidoptera: Lycaenidae) Novitates Zoologicae 34: 374-394. BHL. Reference page.
Sáfián, S. 2017. Three new species in the genus Iolaus Hübner, 1819 (Lepidoptera: Lycaenidae: Theclinae) from West Africa. Metamorphosis 28: 2–10. Full article (PDF). Reference page.
Sharpe, E., 1904. On new species of butterflies from equatorial Africa. Entomologist. 37: 202-204. BHL. Reference page.
Stempffer, H. & Bennett, N.H. 1958. Revision des genres appartenant au groupe des Iolaus (Lep. Lycaenidae). (Premiere Partie). Bulletin de la l’Institut Francais d’Afrique Noire (A) 20: 1243–1347. Reference page.
Stempffer, H. 1967. The genera of the African Lycaenidae (Lepidoptera: Rhopalocera). Bulletin of the British Museum (Natural History), entomology. Supplement 10: 322pp, 1pl. BHL Reference page.
Tshikolovets, V.V., 2011. Butterflies of Europe & the Mediterranean Area Tshikolovets Publications, Pardubice, Czech Republic. ISBN 9788090490000. Reference page.
Williams, M.C. 2007. Checklist of Afrotropical Papilionoidea and Hesperioidea. 7th Edn. (April 2007). Downloaded from Afrotropical Butterflies. Reference page.

Links

Williams, M. Afrotropical Butterflies. Consulted 26.iv.2009.

Iolaus is a genus of butterflies in the family Lycaenidae. The genera Argiolaus, Epamera, Iolaphilus and Stugeta are often included in Iolaus. Iolaus species are found in the Afrotropical realm.There are ca. 130 species in Iolaus
Species
Adalbert Seitz's Fauna Africana pars Iolaus Africa

.

Subgenus Iolaus Hübner, [1819]
Iolaus bilineata Bethune-Baker, 1908
Iolaus bolissus Hewitson, 1873
Iolaus carina Hewitson, 1873
Iolaus eurisus (Cramer, [1780])
Subgenus Epamera Druce, 1891
Iolaus adorabilis Collins & Larsen, 2008
Iolaus aemulus Trimen, 1895
Iolaus aethes Clench, 1965
Iolaus aethria Karsch, 1893
Iolaus agnes Aurivillius, 1898
Iolaus alienus (Trimen, 1898)
Iolaus apatosa (Stempffer, 1952)
Iolaus aphnaeoides Trimen, 1873
Iolaus arborifera (Butler, 1901)
Iolaus aurivillii Röber, 1900
Iolaus australis Stevenson, 1937
Iolaus bakeri (Riley, 1928)
Iolaus bamptoni (Congdon & Collins, 1998)
Iolaus banco Stempffer, 1966
Iolaus bansana Bethune-Baker, 1926
Iolaus belli Hewitson, 1869 (status unclear)
Iolaus bellina (Plötz, 1880)
Iolaus coelestis Bethune-Baker, 1926
Iolaus congdoni (Kielland, 1985)
Iolaus creta Hewitson, 1878
Iolaus cytaeis Hewitson, 1875
Iolaus diametra Karsch, 1895
Iolaus djaloni Collins & Larsen, 1998
Iolaus dubiosa (Stempffer & Bennett, 1959)
Iolaus farquharsoni (Bethune-Baker, 1922)
Iolaus flavilinea (Riley, 1928)
Iolaus fontainei (Stempffer, 1956)
Iolaus frater (Joicey & Talbot, 1921)
Iolaus gemmarius (Druce, 1910)
Iolaus glaucus Butler, 1886
Iolaus handmani (Gifford, 1965)
Iolaus helenae Henning & Henning, 1989
Iolaus hemicyanus Sharpe, 1904
Iolaus iasis Hewitson, 1865
Iolaus jacksoni (Stempffer, 1950)
Iolaus kelle Stempffer, 1967
Iolaus laon Hewitson, 1878
Iolaus leonis (Riley, 1928)
Iolaus longicauda (Stempffer & Bennett, 1959)
Iolaus maesa (Hewitson, 1862)
Iolaus mafugae (Stempffer & Bennett, 1959)
Iolaus mermis (Druce, 1896)
Iolaus mimosae Trimen, 1874
Iolaus mongiro Stempffer, 1969
Iolaus moyambina (Stempffer & Bennett, 1959)
Iolaus nasisii (Riley, 1928)
Iolaus neavei (Druce, 1910)
Iolaus nolaensis (Stempffer, 1951)
Iolaus normani (Larsen, 1986)
Iolaus nursei Butler, 1896
Iolaus obscurus Aurivillius, 1923
Iolaus penningtoni (Stempffer & Bennett, 1959)
Iolaus pollux Aurivillius, 1895
Iolaus pseudofrater Stempffer, 1962
Iolaus pseudopollux Stempffer, 1962
Iolaus sappirus (Druce, 1902)
Iolaus scintillans Aurivillius, 1905
Iolaus sciophilus (Schultze, 1916)
Iolaus sibella (Druce, 1910)
Iolaus sidus Trimen, 1864
Iolaus silanus (Grose-Smith, 1889)
Iolaus stenogrammica (Riley, 1928)
Iolaus sudanicus Aurivillius, 1905
Iolaus tajoraca Walker, 1870
Iolaus umbrosa (Butler, 1886)
Iolaus violacea (Riley, 1928)
Subgenus Aphniolaus Druce, 1902[1]
Iolaus pallene (Wallengren, 1857)
Subgenus Philiolaus Stempffer & Bennett, 1958
Iolaus alcibiades Kirby, 1871
Iolaus calisto (Westwood, 1851)
Iolaus christofferi Collins & Larsen, 2003
Iolaus ismenias (Klug, 1834)
Iolaus laonides Aurivillius, 1898
Iolaus likpe Collins & Larsen, 2003
Iolaus lukabas Druce, 1890
Iolaus mane Collins & Larsen, 2003
Iolaus newporti Larsen, 1994
Iolaus ofere Collins & Larsen, 2008
Iolaus paneperata Druce, 1890
Iolaus parasilanus Rebel, 1914
Iolaus poecilaon (Riley, 1928)
Iolaus theodori Stempffer, 1970
Iolaus vansomereni (Stempffer & Bennett, 1958)
Subgenus Iolaphilus Stempffer & Bennett, 1958
Iolaus aelianus Staudinger, 1891
Iolaus alexanderi Warren-Gash, 2003
Iolaus carolinae Collins & Larsen, 2000
Iolaus gabunica (Riley, 1928)
Iolaus henryi (Stempffer, 1961)
Iolaus icipe Collins & Larsen, 1998
Iolaus iulus Hewitson, 1869
Iolaus jamesoni (Druce, 1891)
Iolaus menas Druce, 1890
Iolaus schultzei Aurivillius, 1905
Iolaus shaba Collins & Larsen, 1995
Iolaus trimeni Wallengren, 1875
Subgenus Argiolaus Druce, 1891
Iolaus aequatorialis (Stempffer & Bennett, 1958)
Iolaus bergeri (Stempffer, 1953)
Iolaus caesareus Aurivillius, 1895
Iolaus cottrelli (Stempffer & Bennett, 1958)
Iolaus crawshayi (Butler, 1901)
Iolaus dianae Heath, 1983
Iolaus iturensis (Joicey & Talbot, 1921)
Iolaus kayonza (Stempffer & Bennett, 1958)
Iolaus lalos (Druce, 1896)
Iolaus manasei (Libert, 1993)
Iolaus maritimus (Stempffer & Bennett, 1958)
Iolaus montana (Kielland, 1978)
Iolaus ndolae (Stempffer & Bennett, 1958)
Iolaus pamae Heath, 1994
Iolaus silarus Druce, 1885
Iolaus silas (Westwood, [1851])
Iolaus stewarti Heath, 1985
Subgenus Pseudiolaus Riley, 1928
Iolaus lulua (Riley, 1944)
Iolaus poultoni (Riley, 1928)
Subgenus Tanuetheira Druce, 1891
Iolaus timon (Fabricius, 1787)
Subgenus Trichiolaus Aurivillius, 1898
Iolaus argentarius Butler, 1879
Iolaus mermeros (Mabille, 1878)

References

Druce, Hamilton H. (1902). "On some new and little-known Butterflies of the Family Lycænidæ from the African, Australian, and Oriental Regions". Proceedings of the Zoological Society of London. 2 (1): 117–118. doi:10.1111/j.1469-7998.1902.tb08224 (inactive 28 February 2022).

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