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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Superfamilia: Curculionoidea

Familia: Curculionidae
Subfamilia: Scolytinae
Tribus: Ipini
Genera: Acanthotomicus – Dendrochilus – IpsOrthotomicusPityogenes – Pityokteines
Name

Ipini Latreille, 1807
References

Latreille, P.A. 1807. Genera crustaceorum et insectorum secundem ordinem naturalem in familias disposita, iconibus exemplisque plurimus explicata. Tomus tertius. Parisiis et Argentorati: A. Koenig, 258 pp. BHL Reference page.
Donald E. Bright, Robert E. Skidmore 2002: A Catalog of Scolytidae and Platypodidae (Coleoptera): Supplement 2 (1995-1999). National Research Council of Canada, Conseil national de recherches Canada. NRC Research Press. 611 pp.
Gao, L. & Cognato, A.I. 2018. Acanthotomicus suncei, a new sweetgum tree pest in China (Coleoptera: Curculionidae<: Scolytinae: Ipini). Zootaxa 4471(3): 595–599. DOI: 10.11646/zootaxa.4471.3.12 Paywall Reference page.

Ipini is a tribe of bark beetles. It is a monophyletic group.[1]

Beetles of this tribe specialize on conifers.[2] Many species, such as the six-spined engraver beetle (Ips sexdentatus) and the European spruce bark beetle (Ips typographus), produce galleries in wood for their eggs and larvae.

Almost all beetles in this tribe have polygynous mating systems with harems of females. An exception is the monogamous Ips latidens.[3]

Genera include:

Acanthotomicus
Ips
Orthotomicus
Pityogenes
Pityokteines
Pseudips

References

Jordal, B., et al. (2008). Secondary structure alignment and direct optimization of 28S rDNA sequences provide limited phylogenetic resolution in bark and ambrosia beetles (Curculionidae: Scolytinae). Zoologica Scripta 37(1), 43–56.
Sequeira, A. S., et al. (2000). Evolutionary assembly of the conifer fauna: distinguishing ancient from recent associations in bark beetles. Proceedings of the Royal Society of London. Series B: Biological Sciences 267(1460), 2359-66.
Reid, M. (1999). Monogamy in the bark beetle Ips latidens: ecological correlates of an unusual mating system. Ecological Entomology 24(1), 89–94.

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