Memphis

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Charaxinae
Tribus: Anaeini
Genus: Memphis
appias Species Group

Species (3): M. appias – M. xenippa – M. xenocles
arginussa Species Group

Species (10): M. arginussa – M. eubaena – M. herbacea – M. lankesteri – M. lemnos – M. neidhoeferi – M. onophis – M. perenna –M. pithyusa – M. smalli
aureola Species Group

Species (4): M. anna – M. aureola – M. dia – M. polyxo
eribotes Species Group

Species (9): M. acidalia – M. beatrix – M. catinka – M. editha – M. hirta – M. laertes – M. leonida – M. otrere – M. pseudiphis
glauce Species Group

Species (1): M. glauce
hedemanni Species Group

Species (3): M. acaudata – M. hedemanni – M. praxias
iphis Species Group

Species (7): M. alberta – M. boliviana – M. cerealia – M. cluvia – M. iphis – M. lorna – M. moeris
moruus Species Group

Species (3): M. moruus – M. oenomais – M. philumena
pasibula Species Group

Species (2): M. falcata – M. pasibula
polycarmes Species Group

Species (17): M. ambrosia – M. anassa – M. aulica – M. basilia – M. cleomestra – M. forreri – M. grandis – M. laura – M. lineata – M. lyceus – M. marylena – M. mora – M. nenia – M. offa – M. phantes – M. polycarmes – M. proserpina
verticordia Species Group

Species (3): M. artacaena – M. perenna – M. verticordia
unknown Species Group

Species (10): M. elisa – M. juliani – M. maria – M. montesino – M. neidhoeferi – M. paulus – M. salinasi – M. viloriae – M. wellingi – M. xenica
Name

Memphis Hübner, [1819]

Type species: Papilio odilia Stoll,
Synonyms

Corycia Hübner, [1825]
Cymatogramma Doubleday, [1849]
Cymatogramma Westwood, [1850]
Euschatzia Grote, 1898
Rydonia Salazar & Constantino, 2001
Annagrapha Salazar & Constantino, 2001

References

Choimet, X., 2009. Nouveaux Charaxinae néotropicaux. Revue des Lépidoptéristes de France 18 (44): 126-129. Reference page.
Costa M., Viloria, A.L., Attal, S. & Orellana, A.M. 2014. Lepidoptera del Pantepui. Parte II. Description de nuevos Nymphalidae (Papilionoidea). Bulletin de la Société entomologique de France 119(1): 39–52. Reference page.
Costa, M., Viloria Petit, Á.L., Attal, S., Neild, A.F.E. Fratell, S.A. & Benmesbah, M. 2019b. Lepidoptera from the Pantepui. Part VIII New Nymphalidae (Charaxinae and Satyrinae) and Riodinidae (Riodininae). Anartia 29: 20-48. Reference page.
Dias, F.M.S., Riley, T.J., Casagrande, M.M. & Mielke, O.H.H. 2021. An overview of “arginussa” species group of Memphis Hübner, [1819], with the description of a new species from Panama (Lepidoptera: Nymphalidae: Charaxinae). Zootaxa 5061(2): 249–270. DOI: 10.11646/zootaxa.5061.2.2. Paywall. Reference page.
Dias, F.M.S.; Casagrande, M.M.; Mielke, O.H.H. 2012: Taxonomic notes on the leafwing butterflies Memphis leonida (Stoll) and Memphis editha (Comstock), stat. nov. (Lepidoptera: Nymphalidae: Charaxinae). Zootaxa 3343: 16–30. Preview Reference page.
Lamas, G., 2004: Atlas of Neotropical Lepidoptera; Checklist: Part 4A; Hesperioidea-Papilionoidea
de la Maza Elvira, R.G. & de la Maza Elvira, J. 2019a. 1.-Descripción de una nueva subespecie de Memphis proserpina (Salvin, 1869) de la Región de Los Tuxtlas, Veracruz, México (Nymphalidae: Charaxinae) 2.-Una nueva subespecie de Parides anchises (Linnaeus, 1758) del Centro Este de Chiapas, México (Papilionidae: Troidini). Revista de la Sociedad Mexicana de Lepidopterología 7(1): 42-76. ResearchGate Reference page.
de la Maza Elvira, R.G. & de la Maza Elvira, J. 2019c. Descripción de una nueva especie y una nueva subespecie del género Memphis Hübner del sureste de México y su relación con las especies del "complejo philumena" (Nymphalidae: Charaxinae). Revista de la Sociedad Mexicana de Lepidopterología 7(1): 17-19. ResearchGate Reference page.
Pyrcz, T.W., 1995: A New Memphis from the Pantepui, Venezuela (Lepidoptera: Nymphaliade: Charaxinae). Tropical Lepidoptera, 6 (1): 11–13. Full article: [1].

Memphis, described by Jacob Hübner in 1819, is a Neotropical nymphalid butterfly genus in the subfamily Charaxinae.

There are 61 species in the genus.[2] All are Neotropical and the undersides of the wings closely resemble dead leaves. In the past Memphis was considered a subgenus of Anaea.

Species

Listed alphabetically within groups:[1]

Species group pasibula:

Memphis pasibula (Doubleday, [1849])
Memphis falcata (Hopffer, 1874)

Species group aureola:

Memphis anna (Staudinger, 1897)
Memphis aureola (Bates, 1866)
Memphis dia (Godman & Salvin, [1884])
Memphis polyxo (H. Druce, 1874)

Species group verticordia:

Memphis artacaena (Hewitson, 1869) – white-patched leafwing[3]
Memphis perenna (Godman & Salvin, [1884]) – big-spotted leafwing[3]
?Memphis pleione (Godart, 1819)
Memphis verticordia (Hübner, 1824)

Species group arginussa:

Memphis arginussa (Geyer, 1832) – mottled leafwing[3]
Memphis herbacea (Butler & H. Druce, 1872) – scarce leafwing[3]
Memphis lemnos (H. Druce, 1877)
Memphis pithyusa (R. Felder, 1869) – blue leafwing, pale-spotted leafwing[3]

Species group hedemanni:

Memphis acaudata (Röber, 1916)
Memphis hedemanni (R. Felder, 1869) – double-banded leafwing[3]
Memphis praxias (Hopffer, 1874)

Species group glauce:

Memphis glauce (C. Felder & R. Felder, 1862)

Species group appias:

Memphis appias (Hübner, [1825])
Memphis xenippa (Hall, 1935)
Memphis xenocles (Westwood, 1850) – corner-spotted leafwing[3]

Species group polycarmes:

Memphis ambrosia (H. Druce, 1874)
Memphis anassa (C. Felder & R. Felder, 1862)
Memphis aulica (Röber, 1916)
Memphis basilia (Stoll, [1780])
Memphis cleomestra (Hewitson, 1869)
Memphis forreri (Godman & Salvin, [1884]) – Guatemalan leafwing[3]
Memphis grandis (H. Druce, 1877)
Memphis laura (H. Druce, 1877)
Memphis lineata (Salvin, 1869)
Memphis lyceus (H. Druce, 1877)
Memphis mora (H. Druce, 1874)
Memphis nenia (H. Druce, 1877)
Memphis offa (H. Druce, 1877)
Memphis phantes (Hopffer, 1874)
Memphis polycarmes (Fabricius, 1775)
Memphis proserpina (Salvin, 1869) – great leafwing[3]

Species group moruus:

Memphis moruus (Fabricius, 1775) – laurel leafwing[3]
Memphis oenomais (Boisduval, 1870) – Boisduval's leafwing[3]
Memphis philumena (Doubleday, [1849]) – orange-striped leafwing[3]

Species group eribotes:

Memphis acidalia (Hübner, [1819])
Memphis beatrix (H. Druce, 1874)
Memphis catinka (H. Druce, 1877)
Memphis hirta (Weymer, 1907)
Memphis laertes (Cramer, [1775])
Memphis leonida (Stoll, [1782])
Memphis otrere (Hübner, 1825)
Memphis pseudiphis (Staudinger, 1887)

Species group iphis:

Memphis alberta (H. Druce, 1876)
Memphis boliviana (H. Druce, 1877)
Memphis cerealia (H. Druce, 1877)
Memphis cluvia (Hopffer, 1874)
Memphis iphis (Latreille, [1813])
Memphis lorna (H. Druce, 1877)
Memphis moeris (C. Felder & R. Felder, [1867])

Ungrouped:

Memphis eurypyle C. Felder & R. Felder, 1862 – pointed leafwing[4]
Memphis juliani Constantino, 1999
Memphis maria Pyrcz & Neild, 1996
Memphis montesino Pyrcz, 1995
Memphis neidhoeferi (Rotger, Escalante, & Coronado, 1965) – wavy-edged leafwing[3]
Memphis salinasi Pyrcz, 1993
Memphis viloriae Pyrcz & Neild, 1996
Memphis wellingi (Miller & Miller, 1976)

References

*Savela, Markku (April 7, 2019). "Memphis Hübner, [1819]". Lepidoptera and Some Other Life Forms. Retrieved January 9, 2020.
Lamas, G. (2004). Atlas of Neotropical Lepidoptera.
Glassberg, Jeffrey. (2007). A Swift Guide to Butterflies of Mexico and Central America. Sunstreak Books Inc. pp.124-127.
Lotts, Kelly & Naberhaus, Thomas (2017). "Pointed Leafwing Memphis eurypyle C. & R. Felder, 1862". Butterflies and Moths of North America. Retrieved January 9, 2020.