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Cladus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Cladus: Pterygota
Cladus: Metapterygota
Cladus: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Myxophaga
Superfamiliae: Lepiceroidea - Sphaeriusoidea
[list of superfamiliae adapted from Kirejtshuk & Poinar (2006: 155-156)]

overview of familiae

Hydroscaphidae - Lepiceridae - Sphaeriusidae - Torridincolidae - †Haplochelidae




* Anton, E.; Beutel, R.G. 2006: On the head morphology of Lepiceridae (Coleoptera: Myxophaga) and the systematic position of the family and suborder. European journal of entomology, 103: 85–95. PDF
* Arce Pérez, R. 1997: Sinopsis del suborden Myxophaga (Coleoptera) de México. Dugesiana, 4(2): 41-50. [in Spanish, with English abstract]
* Beutel, R.G. 1999: Phylogenetic analysis of Myxophaga (Coleoptera) with a redescription of Lepicerus horni (Lepiceridae). Zoologischer Anzeiger, 237: 291–308.
* Beutel, R.G. 2005: 6. Myxophaga Crowson, 1955. Introduction, phylogeny. Pp. 43-44 in: Beutel, R.G. & Leschen, R.A.B. (volume eds.) Coleoptera, Beetles. Volume 1: Morphology and systematics (Archostemata, Adephaga, Myxophaga, Polyphaga partim). In: Kristensen, N.P. & Beutel, R.G. (eds.) Handbook of zoology. A natural history of the phyla of the animal kingdom. Volume IV. Arthropoda: Insecta. Part 38. Berlin, New York: Walter de Gruyter.
* Beutel, R.G.; Maddison, D.R.; Haas, A. 1999: Phylogenetic analysis of Myxophaga (Coleoptera) using larval characters. Systematic entomology, 24: 171-192.
* Kirejtshuk, A.G.; Poinar, G., jr. 2006: Haplochelidae, a new family of Cretaceous beetles (Coleoptera: Myxophaga) from Burmese amber. Proceedings of the Entomological Society of Washington, 108: 155-164. Abstract page scans
* Catalogue of Palearctic Coleoptera. Vol. 1 ed. I. Löbl, & A. Smetana, Apollo Books, Stenstrup, Denmark, 2003 and 2004, p.25 ISBN 87-88757-73-0


* Australian Faunal Directory
* Tree of Life Web Project. 2007. Myxophaga. Version 07 August 2007 (temporary) in The Tree of Life Web Project
* Atlas of beetles of Russia (a project dedicated to the 100th anniversary of G.G. Jacobson's book "Beetles of Russia")

Vernacular names
Česky: řasožraví
English: myxophagans
日本語: ツブミズムシ亜目 (粘食亜目)
Русский: Миксофага
Suomi: Rihmansyöjät
Svenska: Svampätarbaggar


Myxophaga is the second smallest suborder of Coleoptera after Archostemata, consisting of approximately 65 species of small to minute beetles in four families. They have notopleural sutures, although on some families the sutures may be difficult to see.

The members of this suborder are aquatic and semi-aquatic and feed on algae.

Only two myxophagan genera are found in North America: Sphaerius and Hydroscapha.

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Source: Wikispecies: All text is available under the terms of the GNU Free Documentation License