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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Archostemata
Superfamilia: Cupedoidea

Familia: Ommatidae
Subfamilia (3): †Brochocoleinae – Ommatinae – Tetraphalerinae

Genera incertae sedis: †Fuscicupes – †Rhopalomma
Overview of genera

OmmaTetraphalerus – †Pareuryomma – ...
Name

Ommatidae Sharp & Muir, 1912

References
selected references

Ashman, L.G., Oberprieler, R.G. & Slipinski, A. 2015: Rhopalomma stefaniae gen. et sp. n., the first ommatid beetle from the Upper Jurassic in Australia (Coleoptera: Archostemata: Ommatidae). Zootaxa 3980(1): 136–142. DOI: 10.11646/zootaxa.3980.1.8. Preview (PDF) Reference page.
Lawrence, J.F. 1991: Ommatidae (Archostemata) (=Ommadidae, including Tetraphaleridae). P. 298 in: Lawrence, J.F. (coordinator) 1991: Order Coleoptera, in: Stehr, F.W. (ed.) Immature insects, 2 . Kendall/Hunt Publishing Company, Iowa, USA. ISBN 0-8403-4639-5
Tan, J., Wang, Y., Ren, D., Yang, X., 2012: New fossil species of ommatids (Coleoptera: Archostemata) from the Middle Mesozoic of China illuminating the phylogeny of Ommatidae. BMC evolutionary biology 12: 113. DOI: 10.1186/1471-2148-12-113 Open access. Reference page.

Additional references

Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page.
Hangay, G.; Zborowski, P. 2010: A guide to the beetles of Australia. CSIRO Publishing: Collingwood VIC, Australia. ISBN 9780643094871 ISBN 0643094873
Lawrence, J.F. 1999. The Australian Ommatidae (Coleoptera: Archostemata): new species, larva and discussion of relationships. Invertebrate Taxonomy 13: 369–390. DOI: 10.1071/IT99008

Links

Ommatidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD
Tree of Life Web Project. 1995. Ommatidae. Version 01 January 1995 (temporary) in The Tree of Life Web Project
EoL

Vernacular names
中文: 眼甲科


The Ommatidae are a family of beetles in the suborder Archostemata.[1] The Ommatidae are considered the extant beetle family that has most ancestral characteristics.[2] There are only seven extant species, confined to Australia and South America. However, the geographical distribution was much wider during the Mesozoic spanning across Eurasia and Australia, suggesting that they were widespread on Pangea.[3] So far, 28 extinct genera containing over 100 species of these beetles have been described.[4] Three extant genera have been assigned to this family: Omma, Tetraphalerus and Beutelius.[5] The family is considered to be a subfamily of Cupedidae by some authors,[6] but have been found to be more closely related to Micromalthidae in molecular phylogenies.[7] A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia.[8] Due to their rarity, their ecology is obscure, it is likely that their larvae feed on deadwood.[9]
Contents

1 Genera
1.1 Extinct genera
1.2 Extant genera
2 Gallery
3 References
4 External links

Genera

According to Kirejtshuk, 2020[6] and subsequent literature.
Extinct genera

†Allophalerus Kirejtshuk 2020 Early Jurassic-Late Cretaceous (Asia)
†Blapsium Westwood, 1854 Middle Jurassic (Europe)
†Brochocoleus Hong, 1982 Early Jurassic-Late Cretaceous (Eurasia)
†Bukhkalius Kirejtshuk and Jarzembowski 2020 early Late Cretaceous, (Asia)
†Burmocoleus Kirejtshuk 2020 early Late Cretaceous, (Asia)
†Cionocoleus Ren, 1995 Middle Jurassic-Early Cretaceous (Eurasia)
†Cionocups Kirejtshuk 2020 early Late Cretaceous, (Asia)
†Clessidromma Jarzembowski et al. 2017 early Late Cretaceous, (Asia)
†Diluticupes Ren 1995 Middle Jurassic-Early Cretaceous (Eurasia)
†Echinocups Kirejtshuk and Jarzembowski 2020 early Late Cretaceous, (Asia)
†Eurydictyon Ponomarenko, 1969 Early Jurassic (Asia)
†Jarzembowskiops Kirejtshuk 2020 early Late Cretaceous, (Asia)
†Kirejtomma Li & Cai, 2021 early Late Cretaceous, (Asia)
†Liassocupes Zeuner 1962 Early Jurassic (Europe)
†Limnomma Li & Cai in Li et al. 2021, Middle Jurassic (Asia)
†Lithocupes Ponomarenko, 1966 Late Triassic (Asia)
†Miniomma Li, Yamamoto & Cai, 2020, early Late Cretaceous, (Asia)
†Monticupes Ren 1995 Middle Jurassic-Early Cretaceous (Asia)
†Notocupoides Ponomarenko, 1966 Late Triassic (Asia)
†Odontomma Ren et al. 2006 Early Cretaceous (Asia)
†Paraodontomma Yamamoto 2017 early Late Cretaceous, (Asia)
†Pareuryomma Tan et al. 2012 Early Cretaceous (Asia)
†Polyakius Kirejtshuk 2020 early Late Cretaceous, (Asia)
†Rhabdocupes Ponomarenko, 1966 Late Triassic (Asia)
†Rhopalomma Ashman et al. 2015 Late Jurassic (Australia)
†Stegocoleus Jarzembowski and Wang 2016 early Late Cretaceous, (Asia)
†Tetraphalerites Crowson 1962 Eocene (Europe)
†Zygadenia Handlirsch, 1906 Middle Triassic-Late Cretaceous (Eurasia, ?Australia)

Extant genera

Omma Newman, 1839 Late Triassic-Late Cretaceous (Eurasia), recent (Australia)
Tetraphalerus Waterhouse, 1901 Middle Jurassic-Early Cretaceous (Asia) recent (South America)
Beutelius Escalona et al, 2020 Recent, (Australia)

References

"Australian Faunal Directory". Department of the Environment and Energy. Archived from the original on 2011-03-11. Retrieved 2009-11-21.
Hünefeld, Frank; Marvaldi, Adriana E.; Müller, Bianca; Lawrence, John F.; Beutel, Rolf Georg (2011). "The male postabdomen of the "ancestral" archostematan beetle Tetraphalerus bruchi Heller, 1913 (Ommatidae) and its phylogenetic significance". Arthropod Structure & Development. 40 (2): 146–158. doi:10.1016/j.asd.2010.12.003. PMID 21185393.
Yamamoto, Shûhei (2017). "A new genus of Brochocoleini beetle in Upper Cretaceous Burmese amber (Coleoptera: Archostemata: Ommatidae)". Cretaceous Research. 76: 34–39. doi:10.1016/j.cretres.2017.04.008.
Tan, Jingjing; Wang, Yongjie; Ren, Dong; Yang, Xingke (2012). "New fossil species of ommatids (Coleoptera: Archostemata) from the Middle Mesozoic of China illuminating the phylogeny of Ommatidae". BMC Evolutionary Biology. 12: 113. doi:10.1186/1471-2148-12-113. PMC 3518168. PMID 22776212.
"Ommatidae Species List". Joel Hallan’s Biology Catalog. Texas A&M University. Archived from the original on 2012-10-23.
Kirejtshuk, Alexander G. (2020-02-17). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. Bibcode:2020Geosc..10...73K. doi:10.3390/geosciences10020073. ISSN 2076-3263.
McKenna, Duane D.; Shin, Seunggwan; Ahrens, Dirk; Balke, Michael; Beza-Beza, Cristian; Clarke, Dave J.; Donath, Alexander; Escalona, Hermes E.; Friedrich, Frank; Letsch, Harald; Liu, Shanlin (2019-12-03). "The evolution and genomic basis of beetle diversity". Proceedings of the National Academy of Sciences. 116 (49): 24729–24737. doi:10.1073/pnas.1909655116. ISSN 0027-8424. PMC 6900523. PMID 31740605.
Hörnschemeyer, Thomas (July 2009). "The species-level phylogeny of archostematan beetles-where do Micromalthus debilis and Crowsoniella relicta belong?". Systematic Entomology. 34 (3): 533–558. doi:10.1111/j.1365-3113.2009.00476.x. S2CID 84795808.
Adam., Slipinski (2020). "2:Ommatidae". Australian Beetles Volume 2 : Archostemata, Myxophaga, Adephaga, Polyphaga (part). CSIRO Publishing. ISBN 978-0-643-09731-5. OCLC 1127945248.

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