Permocupedidae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: †Protocoleoptera
Superfamilia: †Permocupedoidea

Familia: †Permocupedidae
Genera: Archicupes – Cytocupes – Cytocupoides – Eocupes – Ichthyocupes – Kaltanicupes – ?Kaltanocoleus – Maricoleus – Permocupes – Protocupoides – Tatarocupes – Uralocupes

check: Protocupes

Name

Permocupedidae Martynov, 1933
Synonyms

Stem: Permocuped-
Authority/date: Martynov, 1933
Type genus: Permocupes Martynov, 1933
Stem: Kaltanocole-
Authority/date: Rohdendorf, 1961
Type genus: Kaltanocoleus Rohdendorf, 1961

References

Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page.
Martynov, A.V. 1933: Permskie iskopaemye nasekomye Arkhangel’skogo Kraya. Chast’ II. Setchatokrylye, vislokrylye i zhuki, s prislozheniem opisaniya dvukh novykh zhukov iz Tikhikh Gor [Permian fossil insects from the Arkhangelsk District. Part II. Neuroptera, Megaloptera and Coleoptera with the description of two new beetles from Tikhie Gory]. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 2: 63–96. [in Russian, not seen]
Rohdendorf, B.B. 1961: Nadotryad Coleopteroidea. Zhestkokryloobraznye [Superorder Coleopteroidea] Pp. 393–469 in: Rohdendorf, B.B.; Becker-Migdisova, E.E.; Martynova, O.M.; Sharov, A.G. (eds): Paleozoyskie nasekomye Kuznetskogo Basseyna [Palaeozoic insects of the Kuznetsk Basin]. Trudy Paleontologicheskogo Instituta Akademiya Nauk SSSR, 85: 1–705. [in Russian, not seen]
Shcherbakov, D.E.; Makarkin, V.N.; Aristov, D.S.; Vasilenko, D.V. 2009: Permian insects from the Russky Island, South Primorye. Russian entomological journal, 18 (1): 7–16. PDF

Links

Paleobiology Database
Permocupedidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD

Permocupedidae is a family of Protocoleopteran stem group beetles. They first appeared during the Early Permian, and were one of the dominant groups of beetles during the Middle Permian. They became rare in the Late Permian, with only one species known from the Triassic, Frankencupes ultimus from the Middle Triassic (Anisian) Röt Formation of Germany.[1] They are thought to have been xylophagous (wood eating), which is presumed to be the ancestral ecology of beetles.[2]
Taxonomy

Kirejtshuk (2020) included the following genera in an expanded (sensu lato) family, with the traditional (sensu stricto) Permocupedidae and related Taldycupedidae as subfamilies Permocupedinae and Taldycupedinae respectively.[3]

Subfamily Permocupedinae Martynov, 1932

Afrocupes Geertsema & van den Heever, 1996 South Africa: Western Cape (Kenmoore Farm), Whitehill Formation, Permian, Cisuralian, Sakmarian/Artinskian, −290.1–279.5 Ma.
Afrocupes firmae Geertsema & van den Heever, 1996 (Type species)
Archicupes Rohdendorf, 1961 (syn Palaeocupes Rohdendorf, 1961) Russia, Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma
Archicupes jacobsoni Rohdendorf, 1961 (Type species)
Cytocupoides Ponomarenko, 1969 Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
Cytocupoides longatus Ponomarenko, 1969 (Type species)
Cytocupes Rohdendorf, 1961Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
Cytocupes angustus Rohdendorf, 1961 (Type species)
Eocupes Rohdendorf, 1961Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma).
Eocupes cellulosus Ponomarenko, 1969
Eocupes lukjanovitshi Rohdendorf, 1961 (Type species)
Frankencupes Ponomarenko & Bashkuev, 2018 Germany: Gambach am Main (Lower Franconia), Röt Formation Middle Triassic, Olenekian/Anisian, −247.2–242.0 Ma.
Frankencupes ultimus Ponomarenko & Bashkuev, 2018 (Type species)
Ichthyocupes Rohdendorf, 1961 Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Kemerovo Region (Tykhta River, Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma.
Ichthyocupes kuznetskiensis Ponomarenko, 1969
Ichthyocupes skoki (Rohdendorf, 1961)
Ichthyocupes tyzhnovi Rohdendorf, 1961 (Type species)
Kaltanocoleus Rohdendorf, 1961. Russia: Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
Kaltanocoleus pospelovi Rohdendorf, 1961 (Type species)
Kaltanicupes Rohdendorf, 1961Russia: Kirov Region (Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Orenburg Region (Kargala), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Sweverodvinian), −265.0–259.0 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma. One undescribed species of this genus was recorded from Indian Barakar Formation of the Lower Permian.
Kaltanicupes acutus Ponomarenko, 1963
Kaltanicupes kargalensis Ponomarenko, 1963
Kaltanicupes kitjakensis Ponomarenko, 1963
Kaltanicupes richteri Rohdendorf, 1961 (Type species)
Kaltanicupes reichardti Rohdendorf, 1961
Linicupes Ponomarenko, Yan & Huang, 2014[4] China: Anhui (Houdong, Yinping Mountain, SW Chaohu City), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian/Vyatkian), −265.0–259.0 Ma.
Linicupes yinpinensis Ponomarenko, Yan & Huang, 2014 (Type species)
Maricoleus Shcherbakov in Shcherbakov, Makarkin, Aristov & Vasilenko, 2009 Russia, Primorsky Krai (Cape, Novosilsky, Russky Island), Lower Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
Maricoleus valentinae Shcherbakov in Shcherbakov, Makarkin, Aristov & Vasilenko, 2009 (Type species)
Permocupes Martynov, 1932 (syn Permocupoides Martynov, 1932) Russia: Arkhangel’sk Region (Letopala River and Soyana), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma: Tatarstan (Mendeleevsk District, Tikhie Gory), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
Permocupes distinctus (Ponomarenko, 1963
Permocupes latus Ponomarenko, 2013
Permocupes semenovi Martynov, 1932
Permocupes sojanensis (Ponomarenko, 1963
Pintolla Kirejtshuk, 2020. Brazil: Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Cisuralian, Artinskian/Kungurian (Cisuralian/Biarmian, Kungurian/Kazanian), −279.5–272.5 Ma.
Pintolla ponomarenkoi (Pinto, 1987) (Type species)
Protocupoides Rohdendorf, 1961 (syn Tomiocupes Rohdendorf, 1961, Tricupes Rohdendorf, 1961) Russia: Vologda Region (Isady locality, Mutovino), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinskian/Vyatkian), −259.0–254.0 Ma; Udmurtia (Chepanikha, Rossokha River Valley), Permian, Guadalupian, Wordian (Biarmian, Urzhumian), −268.0–265.0 Ma; Kemerovo Region (Sokolova, Kutsnetsk Basin), Permian, Lopingian, Wuchiapingian/Changhsingian (Tatarian, Severodvinian/Vyatkian), −259.0–252.3 Ma; Kemerovo Region (Kaltan), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
Protocupoides acer (Rohdendorf, 1961)
Protocupoides carinatus (Rohdendorf, 1961)
Protocupoides elongatus Ponomarenko, 2013
Protocupoides esini Ponomarenko, 2013
Protocupoides plavilstshikovi Rohdendorf, 1961
Protocupoides sharoviRohdendorf, 1961
“Permocupes” rohdendorfi Pinto, 1987 Brazil (Rio Grande do Sul (Road Cut on Road BR 290 near Minas do Leao), Irati Formation, Permian, Kungurian, −279.5–272.5 Ma) considered indeterminate to genus by Kirejtshuk (2020)
Tatarocupes Ponomarenko, 2004 Russia: Orenburg Region (Novo-Aleksandrovka), Permian, Guadalupian/Lopingian, Capitanian/Wuchiapingian (Tatarian, Severodvinian), −265.0–259.0 Ma.
Tatarocupes granulatus Ponomarenko, 2004 (Type species)
Uralocupes Ponomarenko, 1969 Russia: Belebeevo Formation Kirov Region (Kityak), Permian, Cisuralian/Guadalupian, Kungurian/Roadian (Cisuralian/Biarmian, Kungurian/Kazanian), −272.5–268.0 Ma.
Uralocupes major Ponomarenko, 1963 (Type species)

Subfamily Taldycupedinae Rohdendorf, 1961

Afrotaldycupes Kirejtshuk, 2020 Beaufort Group, South Africa, Late Permian (Changhsingian)
Afrotaldycupes africanus (Ponomarenko, 2005) (Type species)
Afrotaldycupes lidgettoniensis (Ponomarenko, 2005)
Mesothoris Tillyard, 1916 Blackstone Formation, Australia Late Triassic (Norian)
Mesothoris clathrata Tillyard, 1916 (Type species)
Mesothoris grandis Dunstan, 1923
Mesothoris punctomarginum Dunstan, 1923
Proterocupes Ponomarenko, 2015 (syn Lobanovia Kirejtshuk, 2020)
Proterocupes major Ponomarenko, 2015 Vokhma Formation, Vologda, Russia, Late Permian (Changhsingian)
Proterocupes nedubrovensis Ponomarenko, 2015 Vokhma Formation, Vologda, Russia, Late Permian (Changhsingian)
Proterocupes permiana (Ponomarenko, 2015) (Type species) Poldarsa Formation, Vologda, Russia, Late Permian (Wuchiapingian)
Simmondsia Dunstan, 1923 Blackstone Formation, Australia Late Triassic (Norian)
Simmondsia cylindrica Dunstan, 1923
Simmondsia subpiriformis Dunstan, 1923 (Type species)
Taldycupes Rohdendorf, 1961 (syn Cryptocupes Rohdendorf, 1961, Taldycupidium Rohdendorf, 1961)
Taldycupes bergi (Rohdendorf, 1961) Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
Taldycupes cellulosus Ponomarenko, 2013 Poldarsa Formation, Vologda, Russia, Late Permian (Wuchiapingian)
Taldycupes khalfini Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
Taldycupes moltshanovi Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
Taldycupes pingi Ponomarenko et al. 2014 Yinping Formation, Anhui, China, Middle Permian (Capitanian)
Taldycupes reticulatus Ponomarenko, 1969 Akkolka Formation, Kazakhstan, Late Permian (Changhsingian)
Taldycupes rosanovi Ponomarenko, 2006 Limptekon Formation, Krasnoyarsk, Russia, Late Permian (Changhsingian)
Taldycupes vasjuchitshevi Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
Tecticupes Rohdendorf, 1961 (syn Stegocupes Rohdendorf, 1961
Tecticupes heckeri Rohdendorf, 1961 (Type species) Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian) Mitina Formation, Kemerovo, Russia, Middle Permian (Roadian)
Tecticupes indistinctus Rohdendorf, 1961 Kazankovo-Markinskaya Formation, Kemerovo, Russia, Middle Permian (Wordian)
Tecticupes martynovi Ponomarenko, 2013 Amanak Formation. Orenburg, Middle Permian (Capitanian)
Tychticupes Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
Tychticupes beljanini Rohdendorf, 1961
Tychticupes radtshenkoi Rohdendorf, 1961
Tychticupes ragozini Rohdendorf 1961
"Tychticupes ussovi" Rohdendorf 1961 (nomen dubium)
Tychticupoides Rohdendorf, 1961 Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
Tychticupoides grjazevi Rohdendorf, 1961
Uskaticupes Rohdendorf, 1961 (nomen dubium) Gramoteinskaya Formation, Kemerovo Oblast, Russia, Late Permian
Uskaticupes javorskyi Rohdendorf, 1961 (nomen dubium)

References

Ponomarenko, Alexandr G.; Bashkuev, Alexei S. (December 2018). "First Triassic record of the beetle family Permocupedidae (Insecta: Coleoptera): a peculiar example of a Lazarus taxon". PalZ. 92 (4): 587–591. doi:10.1007/s12542-018-0426-4. ISSN 0031-0220.
Zhao, Xianye; Yu, Yilun; Clapham, Matthew E; Yan, Evgeny; Chen, Jun; Jarzembowski, Edmund A; Zhao, Xiangdong; Wang, Bo (2021-11-08). Perry, George H; Fikacek, Martin (eds.). "Early evolution of beetles regulated by the end-Permian deforestation". eLife. 10: e72692. doi:10.7554/eLife.72692. ISSN 2050-084X.
Kirejtshuk, Alexander G. (2020-02-17). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. Bibcode:2020Geosc..10...73K. doi:10.3390/geosciences10020073. ISSN 2076-3263.
A. G. Ponomarenko, E. V. Yan and Di-ying Huang (2014). "New beetles (Coleoptera) from the terminal Middle Permian of China". Paleontological Journal 48 (2): 191–200. doi:10.1134/S0031030114010109.