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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Nymphalidae
Subfamilia: Satyrinae
Tribus: Haeterini
Genus: Pierella
Species (17): P. amalia – P. angeloi – P. astyoche – P. boliviana – P. chalybaea – P. helvina – P. hortona – P. hyalinus – P. hyceta – P. keithbrowni – P. kesselringi – P. lamia – P. lena – P. lucia – P. luna – P. nereis – P. nice

Pierella Herrich-Schäffer, 1865

Type species: Papilio nereis Drury, [1782]

Pieris Hübner, [1819] (preoccupied by Pieris Schrank, 1801)
Pierella Westwood, [1851] (invalid).
Antirrhaea Westwood, 1851 (unavail.)


Lamas, G. 2004. (ed.) Checklist: Part 4A. Hesperioidea - Papilionoidea. In Heppner, J.B. (ed.) Atlas of Neotropical Lepidoptera. Vol.5A, Pt.4A. Assn. for Tropical Lepidoptera/Scientific Publishers, Gainesville. 439pp. Reference page.
Paluch, M., Zacca, T., Siewert, R.R., Mielke, O.H.H. & Casagrande, M.M. 2015: A new subspecies of Pierella lena (Lepidoptera: Nymphalidae: Satyrinae) in northeastern Brazil. Florida entomologist 98(3): 916–920. Full article (pdf) Reference page.
Salazar, E, J.A. & Constantino, L.M., 2007: New species of (Lepidoptera: Pieridae, Nymphalidae, Satyrinae, Ithomiinae, Riodinidae) in Colombia. Boletín Científico Museo de Historia Natural de Caldas, 11 (1): 167–186. Full article: [1]
Zacca, T., Siewert, R.R., Casagrande, M.M., Mielke, O.H.H. & Paluch, M. 2016. Taxonomic revision of the “Pierella lamia species group” (Lepidoptera: Nymphalidae: Satyrinae) with descriptions of four new species from Brazil: IN Living among dragons: A festschrift in honor to Dr. Angelo B. M. Machado on occasion of his 80th birthday. Zootaxa 4078(1): 366–386. DOI: 10.11646/zootaxa.4078.1.31. Full article (PDF) Reference page.

Pierella is a butterfly genus from the subfamily Satyrinae in the family Nymphalidae found from Mexico through Central America to South America. The species of Pierella have larger hindwings than forewings, unique among butterflies. The oval green flash on the forewing is also unique. It is caused by diffraction, the wing scales forming a diffraction grating.

Their caterpillars have been found on the host plants Heliconia and Calathea.
Species and subspecies

Pierella amalia Weymer, 1885 (= P. lena ab. leucospila)
Pierella astyoche (Erichson, [1849]) – Astyoche satyr
Pierella astyoche astyoche (= Haetera larymna, Hetaera [sic] astyoche)
Pierella astyoche bernhardina Bryk, 1953 (= P. astyoche f. obscura (nomen nudum))
Pierella astyoche stollei Ribeiro, 1931
Pierella helvina (Hewitson, 1859)
Pierella helvina helvina (= Haetera helvina) – red-washed satyr
Pierella helvina hymettia Staudinger, [1886] (= P. incanescens werneri)
Pierella helvina incanescens Godman & Salvin, 1877 (= P. incanescens costaricana)
Pierella helvina ocreata Salvin & Godman, 1868[verification needed] (= P. ocreata (nomen nudum))
Pierella helvina pacifica Niepelt, 1924 (= Pierello [sic] ocreata johnsoni)
Pierella hortona (Hewitson, 1854) – white-barred lady slipper[1]
Pierella hortona hortona (= Haetera hortensia, Haetera hortona, P. hortona f. albopunctata, P. hortona f. ocellata)
Pierella hortona albofasciata Rosenberg & Talbot, 1914 (= P. albofaciata [sic] decepta)
Pierella hyalinus (Gmelin, [1790]) – glassy pierella
Pierella hyalinus hyalinus (= Papilio hyalinus, Pierella hyalinus fusimaculata, Pieris dracontis)
Pierella hyalinus extincta Weymer, 1910
Pierella hyalinus schmidti Constantino, 1995
Pierella hyalinus velezi Constantino, 1995
Pierella hyceta (Hewitson, 1859) – golden lady slipper[2]
Pierella hyceta hyceta (= Haetera hyceta)
Pierella hyceta ceryce (Hewitson, 1874) (= Haetera ceryce)
Pierella hyceta latona (C.Felder & R.Felder, 1867) (= Haetera latona)
Pierella incanescens (Godman & Salvin, 1877)
Pierella lamia (Sulzer, 1776) – Sulzer's lady slipper[3]
Pierella lamia lamia (= Papilio dyndimene, Papilio lamia, Papilio rhea, Pierella lamia f. fabriciana , Pierella luna ab. albina)
Pierella lamia boliviana F.M.Brown, 1948
Pierella lamia chalybaea Godman, 1905 (= Haetera lamia f. columbina)
Pierella lena (Linnaeus, 1767) – Lena pierella
Pierella lena lena (= Papilio lena, Papilio sectator)
Pierella lena brasiliensis (C.Felder & R.Felder, 1862) (= Haetera brasiliensis, P. lena browni, P. lena glaucolena, P. lena f. melanosa (nomen nudum), P. lena f. obsoleta)
Pierella lucia Weymer, 1885 (= P. astyoche var. albomaculata) – Lucia pierella
Pierella luna (Fabricius, 1793) – Moon Satyr
Pierella luna luna (= Papilio luna, Pierella luna f. rubra)
Pierella luna lesbia Staudinger, 1887
Pierella luna pallida (Salvin & Godman, 1868) (= Hetaera [sic] pallida)
Pierella luna rubecula Salvin & Godman, 1868[verification needed] (= Haetera heracles)
Pierella nereis (Drury, 1782) (= Papilio nereis)[4]


White-barred Lady Slipper, Learn about Butterflies
Golden Lady Slipper, Learn about Butterflies
Sulzer's Lady Slipper, Learn about Butterflies

"Pierella nereis". iNaturalist.ca. Retrieved 2021-06-03.

Nymphalidae Study Group species list
Urich, F. C., and T. C. Emmel. 1990. Life histories of Neotropical butterflies from Trinidad 1. Pierella hyalinus fusimaculata (Lepidoptera: Satyridae). Tropical Lepidoptera 1(1): 25-26.pdf

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