Progonocimicidae

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Paraneoptera
Superordo: Condylognatha
Ordo: Hemiptera
Subordo: Coleorrhyncha

Familia: †Progonocimicidae
Genera (21): †Progonocimex – †Actinoscytina – †Archicercopis – †Cicadocoris – †Eocercopis – †Heterojassus – †Heteroscytina – †Hexascytina – †Ildavia – †Ilahulgabalus – †Mesoscytina – †Microscytinella – †Olgamartynovia – †Platyscytinella – †Popovigocimex – †Popovus – †Triassocoecus – †Triscytina – †Valdiscytina – †Yurigocimex – †Yuripopovia

Name

Progonocimicidae Handlirsch, 1906

Bibliographic reference: Handlirsch, 1906, p. 493.

Type genus: Progonocimex Handlirsch, 1906 (=Eocimex Handlirsch, 1906)
Synonyms

Eocimicidae Handlirsch, 1906 (p. 494)
Actinoscytinidae J.W. Evans, 1956 (p. 244)
Cicadocoridae Becker-Migdisova, 1958 (p. 60)
Actinocytinidae Carpenter, 1992 (p. 259) lapsus calami

References

Becker-Migdisova, E.E. 1958. New fossil Hemiptera. Materialy k Osnovam Paleontologii 2: 57–67.
Carpenter, F.M. 1992. Superclass Hexapoda. Volumes 3 & 4, Part R, in Kaesler, R.L. (ed.) Treatise on Invertebrate Paleontology. Geological Society of America, Boulder, Colorado & University of Kansas, Lawrence, Kansas, 655 pp.
Dong, Q., Yao, Y. & Ren, D. 2014. New fossil Progonocimicidae (Hemiptera: Coleorrhyncha: Progonocimicoidea) from the Upper Mesozoic of northeastern China, with a phylogeny of Coleorrhyncha. Systematic entomology 39(4): 773–782. DOI: 10.1111/syen.12085 Reference page.
Evans, J.W. 1956. Palaeozoic and Mesozoic Hemiptera (Insecta). Australian Journal of Zoology 4: 165–258.
Handlirsch, A. 1906–1908. Die Fossilien Insecten und die Phylogenie der Rezenten Formen; ein Handbuch für Paläontologen und Zoologen. Engelmann, Leipzig, 1430 pp.
Szwedo, J., Azar, D. & Ziadé, K. 2011. The first Progonocimicidae (Insecta: Hemiptera: Coleorrhyncha) from Lower Cretaceous Lebanese amber. Insect systematics & evolution 42(2): 161–177. DOI: 10.1163/187631211X578415

The Progonocimicidae are an extinct family of true bugs in the suborder Coleorrhyncha. Progonocimicidae fossils have been found in Europe, Asia, Australia, and South America.[1][2]

Families preserved in the fossil record of Coleorrhyncha include the early Progonocimicidae, and the later Karabasiidae and Hoploridiidae. The only extant family in Coleorrhyncha is Peloridiidae.[2][3][4] The family is thought to have evolved from the Permian Ingruidae.[5]

Genera

These 27 genera belong to the family Progonocimicidae:[1]

†subfamily Cicadocorinae Bekker-Migdisova 1958
† Absoluta Becker-Migdisova, 1962 Dzhil Formation, Kyrgyzstan, Hettangian
† Archicercopis Handlirsch, 1939 Green Series, Germany, Toarcian
† Cicadocoris Becker-Migdisova, 1958 Dzhil Formation, Kyrgyzstan, Hettangian Abasheva Formation, Russia, Pliensbachian, Sagul Formation, Kyrgyzstan, Toarcian, Itat Formation, Russia, Bajocian/Bathonian Daohugou, China, Callovian Ichetuy Formation, Russia, Oxfordian, Kalgan Formation, Russia, Kimmeridgian
†Gakasha Jiang et al. 2018 Burmese amber, Myanmar, Cenomanian
† Ilahulgabalus Szwedo, Azar & Ziade, 2011 Lebanese amber, Barremian
† Ildavia Popov, 1993 Weald Clay, United Kingdom, Hauterivian
†Mesocimex Hong 1983 Daohugou, Haifanggou Formation, China, Callovian
†Olgamartynovia Becker-Migdisova 1958 Dzhil Formation, Kyrgyzstan, Hettangian Abasheva Formation, Russia, Pliensbachian, Sagul Formation, Kyrgyzstan, Toarcian, Itat Formation, Russia, Bajocian/Bathonian Daohugou, China, Callovian Ichetuy Formation, Russia, Oxfordian, Kalgan Formation, Russia, Kimmeridgian
† Onokhoia Popov, 1988 Godymboyskaya Formation, Russia, Aptian
† Valdiscytina Popov, 1993 Weald Clay, United Kingdom, Hauterivian
† Yuripopovia Jarzembowski, 1991 Weald Clay, United Kingdom, Hauterivian, Vectis Formation, United Kingdom, Aptian
† Actinoscytina Tillyard, 1926 Croudace Bay Formation, Australia, Changhsingian
† Eocercopis Handlirsch, 1939 Green Series, Germany, Toarcian
† Heterojassus Evans, 1961 Mount Crosby Insect Bed, Australia, Norian
† Heteronella Evans, 1961 Mount Crosby Insect Bed, Australia, Norian
† Heteroscytina Evans, 1956
† Hexascytina Wootton, 1963 Mount Crosby Insect Bed, Australia, Norian
† Indutionomarus Szwedo, 2011 Bascharage, Luxembourg, Toarcian
† Microscytinella Wootton, 1963 Mount Crosby Insect Bed, Australia, Norian
† Ovicimex Hong & Wang, 1990 Laiyang Formation, China, Aptian
† Pelorisca Popov & Shcherbakov, 1991 Madygen Formation, Kyrgyzstan, Ladinian
† Platyscytinella Evans, 1956 Mount Crosby Insect Bed, Australia, Norian
† Popovigocimex Martins-Neto & Galego, 2003 Los Rastros Formation, Argentina, Carnian
† Popovus Özdikmen & Demir, 2007 Gurvan-Eren Formation, Mongolia, Aptian
† Progonocimex Handlirsch, 1906 Green Series, Germany, Toarcian
†Pseudipsvicia Handlirsch 1939 Blackstone Formation, Australia, Norian
† Triassodoecus Evans, 1963 Hawkesbury Sandstone, Australia, Anisian
† Triscytina Evans, 1956 Mount Crosby Insect Bed, Australia, Norian
† Woottonia Popov & Shcherbakov, 1991 Tologoi Formation, Kazakhstan, Norian
† Yurigocimex Martins-Neto & Galego, 2003 Los Rastros Formation, Argentina, Carnian

References

Heads, Sam W.; Maehr, Michael D.; Eades, David C. (2019). "family Progonocimicidae Handlirsch, 1906". Coleorrhyncha species file online, Version 5.0. Retrieved 2019-05-03.
Progonocimicidae at Fossilworks
A new species of Progonocimicidae (Hemiptera, Coleorrhyncha) from the Middle Jurassic of China
Heads, Sam W. (2008). "A new species of Yuripopovia (Coleorrhyncha: Progonocimicidae) from the Early Cretaceous of the Isle of Wight" (PDF). British Journal of Entomology and Natural History. 21: 247–253.
Jiang, Tian; Wang, Bo; Szwedo, Jacek (January 2019). "The first representative of Progonocimicidae (Hemiptera: Coleorrhyncha) from mid-Cretaceous Burmese amber". Cretaceous Research. 93: 346–359. doi:10.1016/j.cretres.2018.09.018.