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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Orthoptera
Subordo: Ensifera
Infraordo: Tettigoniidea
Superfamilia: Tettigonioidea

Familia: Tettigoniidae
Subfamilia: Pseudophyllinae
Tribus: Aphractini - Aspidonotini - Callimenellini - Cocconotini - Cymatomerini - Eucocconotini - Homalaspidiini - Ischnomelini - Leptotettigini - Pantecphylini - Phyllomimini - Platyphyllini - Pleminiini - Polyancistrini - Pseudophyllini - Pseudotomias – Pterochrozini - Pterophyllini - Simoderini - Teleutiini - Unassigned
Name

Pseudophyllinae Burmeister, 1840
References

Hemp, C. 2016. The Eastern Arc Mountains and coastal forests of East Africa—an archive to understand large-scale biogeographical patterns: Pseudotomias, a new genus of African Pseudophyllinae (Orthoptera: Tettigoniidae). Zootaxa 4126(4): 480–490. DOI: 10.11646/zootaxa.4126.4.2. Reference page.
Hugel, S.; Desutter-Grandcolas, L. 2013: New Pseudophyllinae from the Lesser Antilles (Orthoptera: Ensifera: Tettigoniidae). Zootaxa 3741(2): 279–288. DOI: 10.11646/zootaxa.3741.2.6 Reference page.

Vernacular names
日本語: ヒラタツユムシ亜科

The subfamily Pseudophyllinae contains numerous species in the family Tettigoniidae, the katydids or bush crickets. Sometimes called "true katydids" , together with the crickets of suborder Ensifera, they form part of the insect order Orthoptera which also contains grasshoppers.

Members of the group are noted for their remarkable camouflage. They closely resemble dried leaves, including veins, various blotches and even bite marks.[1]

Systematics

The Pseudophyllinae may be subdivided into the following tribes (the first 17 of which are sometimes grouped into the super-tribes: Pleminiiti and Pseudophylliti) and genera.[2] Some notable species are also listed here:
Aphractini

Auth.: (Brunner von Wattenwyl 1895) - Chile

Aphractus Redtenbacher, 1895
Paraphractus Beier, 1962
Polycleptidella Beier, 1962
Polycleptis Karsch, 1891

Aspidonotini

Auth.: (Brunner von Wattenwyl 1895) - Madagascar
Aspidonotus spinosus

Aspidonotus Brullé, 1835

Callimenellini

Auth.: (Gorochov 1990) - Asia: including India, Indo-China and the Philippines

Callimenellus Walker, 1871

Cocconotini

Auth.: (Brunner von Wattenwyl 1895) - Central and southern America

Beierotettix Özdikmen, 2007
Bliastes Stål, 1873
Bliastonotideus Beier, 1962
Bliastonotus Beier, 1960
Brachybliastes Beier, 1960
Calamoptera Saussure, 1861
Chrysobliastes Beier, 1960
Cobalotettix Beier, 1962
Cocconotus Stål, 1873
Cojedebius Kevan, 1989
Diplopygia Beier, 1962
Docidocercus Beier, 1960
Eubliastes Beier, 1960
Geonotus Beier, 1960
Helicocercus Beier, 1960
Idiarthron Brunner von Wattenwyl, 1895
Incanotus Beier, 1960
Jamaicoecia Beier, 1962
Leiobliastes Beier, 1960
Liparoscelis Stål, 1873
Liparoscella Hebard, 1933
Melanonotus Brunner von Wattenwyl, 1895
Meroncidius Serville, 1831
Mystron Montealegre-Z. & Morris, 1999
Nannonotus Beier, 1960
Nannotettix Redtenbacher, 1895
Nastonotus Bolívar, 1890
Natagaima Beier, 1960
Nesoecia Scudder, 1893
Nesonotus Beier, 1960
Schedocentrus Hebard, 1924
Sphaeropyga Beier, 1960
Stenotettix Stål, 1873
Thamnobates Saussure & Pictet, 1898
Trichotettix Stål, 1873

Cymatomerini

Auth.: (Brunner von Wattenwyl 1895) - Central Africa and Asia: including India, Indo-China, Malesia and the Philippines
Sanaa regalis
Tegra novaehollandiae

Cymatomera Schaum, 1853 (Africa)
Cymatomerella Beier, 1954 (Africa)
Olcinia Stål, 1877
Parasanaa Beier, 1944
Platenia Dohrn, 1888 - monotypic Platenia semialata (Philippines)
Sanaa Walker, 1870
Sathrophyllia Stål, 1874
Sathrophylliopsis de Jong, 1939 (W. Malesia)
Tegra Walker, 1870
Tegrolcinia de Jong, 1939 (China, W. Malesia)
Typhoptera Kirby, 1906

Eucocconotini

Auth.: Beier 1960 - Central and South America
Panoploscelis specularis

Dicranostomus Dohrn, 1888
Eucocconotideus Beier, 1960
Eucocconotus Hebard, 1927
Gnathoclita Hagenbach, 1842
Myopophyllum Beier, 1960
Onychopygia Beier, 1962
Panoploscelis Scudder, 1869

Homalaspidiini

Auth.: (Otte, D. 1997) - Southern America

Arrhenotettix Caudell, 1918
Disceratus Scudder, 1869
Homalaspidia Uvarov, 1940
Jimenezia Bolívar, 1881
Sterphoter Rehn, 1946

Ischnomelini

Auth.: (Beier 1960) - Southern America

Goethalsiella Hebard, 1927
Ischnomela Stål, 1873

Leptotettigini

Auth.: (Beier 1960) - Southern America

Chondrosternum Beier, 1960
Leptotettix Stål, 1873
Macrochiton Redtenbacher, 1895
Neochiton Beier, 1962
Pezochiton Beier, 1960
Platychiton Beier, 1960
Semileptotettix Brunner von Wattenwyl, 1895

Pantecphylini

Auth.: (Brunner von Wattenwyl 1895) - Central Africa

Pantecphylus Karsch, 1890

Phrictini

Auth.: Bolívar, 1903 - sometimes placed in the Mecopodinae (Eastern Australia)
Phricta aberrans

Phricta Redtenbacher, 1892

Phyllomimini

Auth.: (Brunner von Wattenwyl 1895) - West and central Africa; Australasia: India through to Pacific islands

Acanthoprion Pictet & Saussure, 1892
Acauloplacella Karny, 1931
Acauloplax Karsch, 1891
Brochopeplus Pictet & Saussure, 1892
Brunneriana Uvarov, 1923
Chondrodera Karsch, 1890
Chondroderella Hebard, 1922
Despoina Brunner von Wattenwyl, 1895
Diplodontopus Karny, 1931
Gonyatopus Brunner von Wattenwyl, 1895
Hapalophyllum Hebard, 1922
Hemigyrus Brunner von Wattenwyl, 1893
Heteraprium Krauss, 1903
Lacipoda Brunner von Wattenwyl, 1895
Lagarodes Karsch, 1890
Margarodera Beier, 1957
Mioacris Pictet & Saussure, 1892
Morsimus Stål, 1877
Orophyllus Beier, 1954
Paramorsimus Beier, 1954
Paraphyllomimus Beier, 1954
Phyllomimus Stål, 1873
Phyllozelus Redtenbacher, 1892
Pirmeda Henry, 1940
Promeca Brunner von Wattenwyl, 1895
Pseudacanthoprion Beier, 1954
Pseudophyllomimus Karny, 1924
Rhinodera Beier, 1955
Stenampyx Karsch, 1890
Stizoscepa Karsch, 1896
Temnophylloides Henry, 1939
Temnophyllus Redtenbacher, 1895
Timanthes Stål, 1877
Tomias Karsch, 1890
Tympanophyllum Krauss, 1903
Tympanoptera Redtenbacher, 1892
Zumala Walker, 1869

Platyphyllini

Auth.: (Brunner von Wattenwyl 1895) - Southern America

Acyrophyllum Beier, 1960
Aemasia Brunner von Wattenwyl, 1895
Antillophyllum Beier, 1960
Baliophyllum Beier, 1962
Brachyauchenus Brunner von Wattenwyl, 1895
Choeroparnops Dohrn, 1888
Diyllus Stål, 1875
Drepanoxiphus Brunner von Wattenwyl, 1895
Haenschiella Beier, 1960
Kopis Beier, 1960
Myrmeciophyllum Beier, 1960
Opetiocercus Beier, 1960
Platyphyllum Serville, 1831
Rhabdotophyllum Beier, 1960
Triencentrus Brunner von Wattenwyl, 1895
Xiphophyllum Beier, 1960

Pleminiini

Auth.: (Brunner von Wattenwyl 1895) - Central and southern America, West and central Africa
Batodromeus subulo

Acanthodis Serville, 1831
Acanthorhinischia Beier, 1954
Adapantus Karsch, 1891
Adeclus Brunner von Wattenwyl, 1895
Adenes Karsch, 1891
Amiltonia Piza, 1976
Ancistrocercus Beier, 1954
Anonistus Walker, 1871
Apereisis Walker, 1871
Aspidopygia Beier, 1962
Balboana Uvarov, 1939
Batodromeus Karsch, 1896
Beieroschema Özdikmen, 2008 (formerly Colobotettix Beier, 1960 (non Ribaut, 1948: preoccupied))
Bufotettix Caudell, 1918
Championica Saussure & Pictet, 1898
Clepsydronotus Beier, 1954
Condylocnemis Redtenbacher, 1895
Corynecercus Beier, 1954
Dasyscelidius Beier, 1954
Dasyscelus Redtenbacher, 1895
Diacanthodis Walker, 1870
Enthacanthodes Redtenbacher, 1895
Gongrocnemis Redtenbacher, 1895
Habrocomes Karsch, 1890
Haemodiasma Brunner von Wattenwyl, 1895
Hoplidostylus Karsch, 1893
Jamaicana Brunner von Wattenwyl, 1895
Labidocercus Beier, 1954
Leurophyllidium Beier, 1954
Leurophyllum Kirby, 1906
Lichenochrus Karsch, 1890
Mormotus Karsch, 1890
Nesokatoikos Braun & Maehr, 2008 (formerly Nesocnemis Beier, 1954 (non Sélys, 1891: preoccupied))
Nesophyllidium Beier, 1954
Orpacanthophora Beier, 1954
Paradeclus Beier, 1954
Paralichenochrus Beier, 1954
Parapleminia Beier, 1954
Pedinothorax Beier, 1954
Phyllostachydius Beier, 1954
Pleminia Stål, 1874
Polyglochin Karsch, 1890
Pristonotus Uvarov, 1940
Pseudopleminia Beier, 1954
Rhinischia Beier, 1954
Schochia Brunner von Wattenwyl, 1895
Sphyrophyllum Beier, 1954
Stenoschema Redtenbacher, 1895
Tetragonomera Stål, 1874
Tympanocompus Karsch, 1891
Xerophyllopteryx Rehn, 1905

Polyancistrini

Auth.: (Brunner von Wattenwyl 1895) - Mexico, Caribbean, Southern America

Camposiella Hebard, 1924
Polyancistroides Rehn, 1901
Polyancistrus Serville, 1831
Sagephorus Redtenbacher, 1895
Spelaeala Rehn, 1943
Spinapecta Naskrecki & Lopes-Andrade, 2005

Pseudophyllini

Auth.: (Burmeister 1838) - West and central Africa; Asia: India, Indo-China through to Papua New Guinea
Onomarchus kanara
Zabalius cf. ophthalmicus

Brunneana Uvarov, 1939
Chloracris Pictet & Saussure, 1892
Climacoptera Redtenbacher, 1895
Cratioma Bolívar, 1906
Desaulcya Bolívar, 1906
Liocentrum Karsch, 1890
Micta Karsch, 1896
Mustius Stål, 1874
Onomarchus Stål, 1874
Opisthodicrus Karsch, 1890
Oxyaspis Brunner von Wattenwyl, 1895
Perteus Bolívar, 1906
Pseudophyllus Serville, 1831
Pseudophyllus titan
Rhomboptera Redtenbacher, 1895
Zabalius Bolívar, 1886
Zabalius apicalis

Pterophyllini

Auth.: (Karny, 1925) – Americas

Caloxiphus Saussure & Pictet, 1898
Caribophyllum Rehn, 1947
Diophanes Stål, 1875
Elytraspis Beier, 1962
Karukerana Bonfils, 1965
Lea Caudell, 1906
Lophaspis Redtenbacher, 1895
Mastophyllum Beier, 1960
Paracyrtophyllus Caudell, 1906
Parascopioricus Beier, 1960
Phyllopectis Rehn, 1948
Pterophylla Kirby, 1825
Scopioricus Uvarov, 1940
Scopiorinus Beier, 1960
Thliboscelus Serville, 1838
Xestoptera Redtenbacher, 1895

Simoderini

Auth.: (Brunner von Wattenwyl, 1895) - Madagascar, Australia

Chloracantha Hebard, 1922
Lonchitophyllum Brunner von Wattenwyl, 1895
Mastigaphoides Weidner, 1965
Mastighapha Karsch, 1891
Parasimodera Carl, 1914
Phyrama Karsch, 1889
Simodera Karsch, 1891
Wattenwyliella Carl, 1914

Teleutiini

Auth.: (Beier, 1960) - Panama to Southern America

Acanthodiphrus Walker, 1871
Apteroteleutias Beier, 1962
Beierella Piza, 1978
Brachyteleutias Beier, 1960
Chibchella Hebard, 1927
Eumecopterus Beier, 1960
Hoplotettix Caudell, 1918
Leptoteleutias Beier, 1962
Mylothrella Beier, 1960
Pemba Walker, 1870
Stetharasa Montealegre-Z. & Morris, 1999
Teleutias Stål, 1874

Tribe undetermined

Narea Walker, 1870 - Australia
†Archepseudophylla Nel, Prokop & Ross, 2008

Due to the great diversity of this subfamily, it is not unequivocally delimited yet.
References

Grimaldi, David; Engel, Michael S. (2005). "Orthoptera: the crickets, katydids, grasshoppers, wetas, and kin". Evolution of the Insects. Cambridge: Cambridge University Press. pp. 202–211. ISBN 0521821495.
Orthoptera Species File (retrieved 25 December 2017)

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