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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Hesperiidae
Subfamilia: Pyrginae
Tribus: Pyrgini
Genus: Pyrgus
Species: P. accreta – P. aladaghensis – P. alpinus – P. alveus – P. andromedae – P. armoricanus – P. badachschana – P. bellieri – P. bolkariensis – P. bieti – P. cacaliae – P. carlinae – P. carthami – P. cashmirensis – P. centaureae – P. centralasiae – P. cinarae – P. darwazica – P. dejeani – P. fides – P. jupei – P. maculatus – P. malvae – P. melotis – P. nepalensis – P. numida – P. oberthueri – P. onopordi – P. ruralis – P. scriptura – P. serratulae – P. sibirica – P. sidae – P. speyeri – P. trebevicensis – P. warrenensis – P. xanthus

Pyrgus Hübner, [1819].

Type species: Papilio alveolus Hübner, [1800-1803], by subsequent designation by Westwood in Humphreys & Westwood, 1841: 120.


Pyrgus sp.


=♦ Urbanus Hübner, [1806] (suppr. ICZN Op. 97); TS: malvae Linnaeus, 1758.
♦ Pyrgus Hübner, [1819]: 109; Humphreys & Westwood, 1841: 119; Hemming, 1967: 389; Bridges, 1994: IX.54.
♦ Scelotrix Rambur, 1858ː 63. TS: carthami Hübner, [1813]
♦ Bremeria Tutt, 1906ː 296. (Invalid; preocc. Bremeria Alpheraky, 1892 Nymphalidae).TS: bieti Oberthür, 1886
♦ Ateleomorpha Warren, 1926: 19, 87; typus onopordi Rambur, 1840.
♦ Hemiteleomorpha Warren, 1926: 19, 72; typus malvae (Linnaeus).
♦ Teleomorpha Warren, 1926: 18, 46; typus carthami (Hübner).
Primary references

Hübner, J. 1816–1826. Verzeichniss bekannter Schmettlinge. Augsburg. Papilionoidea. BHL Reference page.

Additional references

Alberti, B., 1967. Über die Hesperia alveus Hbn. - Gruppe im Kaukasus - Raum nebst Beschreibung einer nellen Art (Lep., Hesperiidae). Deutsche Entomologische Zeitschrift 14: 461-472, 3 figs, 1 map. Reference page.
Bridges, C.A. 1994. Catalogue of the Family-Group, Genus-Group and Species-Group Names of the Hesperioidea (Lepidoptera) of the World. Bridges, Urbana, Illinois. BHL Reference page.
Hemming, A. F., 1967. The generic names of the butterflies and their type species (Lepidoptera: Rhopalocera). Bulletin of the British Museum (Natural History) 1967, Suppl. 9: 1-509.Reference page.
Hernández-Roldán, J.L., Munguira, M.L., Wagner, W. & Vila, R. 2012. Comparative analysis and taxonomic use of the morphology of immature stages and natural history traits in European species of Pyrgus Hübner (Lepidoptera: Hesperiidae, Pyrginae). Zootaxa 3470: 1–71. Preview Reference page.
Humphreys, H.N. & Westwood, J.O. 1841. British Butterflies and Their Transformations. London: William Smith. 138pp, 42 Pls. Reference page.
de Jong, R. 1975. Notes on the genus Pyrgus (Lepidoptera, Hesperiidae). Zoologische Mededelingen 49(1): 1–18. PDF. Reference page.
de Jong, R. 1979. Revision of the Pyrgus alpinus group (Lepidoptera, Hesperiidae), with notes on phylogeny and character displacement. Zoologische Mededelingen 54(4): 53–82. PDF. Reference page.
Korb, S.K. 2015b. Butterflies (Lepidoptera: Papilionoformеs) of the Inner Tian-Shan. Eversmannia Supplement 6: 3-?. Reference page.
Li, W., Cong. Q., Shen, J.H., Zhang, J., Hallwachs, W., Janzen, D.H. & Grishin, N.V. 2019. Genomes of skipper butterflies reveal extensive convergence of wing patterns. PNAS 116(13): 6232–6237. DOI: 10.1073/pnas.1821304116. Text. SI Appendix. Reference page.
Rambur, J. P. 1858–1866. Catalogue systematique des Lepidopteres de l'Andalousie. Paris. J. B. Balliere, 412pp, pls 1–22. Reference page.
Schilde, 1886. Betrachtungen über die Variabilität in der Schmetterlings-Gattung Pyrgus. Berliner Entomologische Zeitschrift 30: 39-62. Reference page.
Tshikolovets, V.V., 2011. Butterflies of Europe & the Mediterranean Area Tshikolovets Publications, Pardubice, Czech Republic. ISBN 9788090490000. Reference page.
Wallengren, 1853. Skandinaviens Hesperiidae med särskild hähsyn till arteme af slägtet Syrichtus. Öfversigt af Kongl. Vetenskaps-akademiens forhandlingar. 10: 19-26. Reference page.
Warren, B.C.S., 1926. Monograph of the Tribe Hesperiidi (European Species) with Revised Classification of the Subfamily Hesperiinae (Palaearctic Species) Based on the Genital Armature of the Males. Transactions of the Entomological Society of London 74(1): 1–167. DOI: 10.1111/j.1365-2311.1926.tb01184.x Reference page.


Vernacular names
English: Grizzled skippers

Pyrgus is a genus in the skippers butterfly family, Hesperiidae, known as the grizzled skippers. The name "checkered" or "chequered skipper" may also be applied to some species, but also refers to species in the genus Carterocephalus. They occur in the Holarctic with an additional group of species extending to the Neotropic.

In 2019, most of the species of Pyrgus found in the North, Central, or South America were moved to the genera Burnsius, Chirgus, and Heliopetes. The remaining Pyrgus species found in the New World are Pyrgus centaureae, ruralis, scriptura, and xanthus.[1]

These species belong to the genus Pyrgus:[1][2]

Pyrgus accretus (Verity, 1925)
Pyrgus alpinus Erschoff, 1874
Pyrgus alveus (Hübner, 1803) (large grizzled skipper)
Pyrgus andromedae (Wallengren, 1853) (Alpine grizzled skipper)
Pyrgus armoricanus (Oberthur, 1910) (oberthür's grizzled skipper)
Pyrgus cacaliae (Rambur, 1839) (dusky grizzled skipper)
Pyrgus carlinae (Rambur, [1839]) (carline skipper)
Pyrgus carthami (Hübner, [1813]) (safflower skipper)
Pyrgus centaureae (Rambur, 1842) (northern grizzled skipper)
Pyrgus centralitaliae (Verity, 1920)
Pyrgus cinarae (Rambur, 1839) (sandy grizzled skipper)
Pyrgus cirsii (Rambur, 1839) (cinquefoil skipper)
Pyrgus foulquieri (Oberthür, 1910) (foulquier's grizzled skipper)
Pyrgus jupei (Alberti, 1967) (caucasian skipper)
Pyrgus maculatus Bremer & Grey, 1853
Pyrgus malvae (Linnaeus, 1758) (grizzled skipper)
Pyrgus malvoides (Elwes & Edwards, 1897) (southern grizzled skipper)
Pyrgus melotis Duponchel, 1832 (aegean skipper)
Pyrgus nepalensis Higgins, 1984
Pyrgus onopordi (Rambur, 1839) (rosy grizzled skipper)
Pyrgus pontica Reverdin, 1914
Pyrgus ruralis (Boisduval, 1852) (two-banded checkered skipper)
Pyrgus scriptura (Boisduval, 1852) (small checkered skipper)
Pyrgus serratulae (Rambur, [1839]) (olive skipper)
Pyrgus sidae (Esper, 1784) (yellow-banded skipper)
Pyrgus speyeri (Staudinger, 1887)
Pyrgus warrenensis (Verity, 1928) (warren's skipper)
Pyrgus xanthus W. H. Edwards, 1878 (mountain checkered skipper)

References and external links

Li, Wenlin; Cong, Qian; Shen, Jinhui; Zhang, Jing; et al. (2019). "Genomes of skipper butterflies reveal extensive convergence of wing patterns". Proceedings of the National Academy of Sciences. 116 (13): 6232–6237. doi:10.1073/pnas.1821304116. PMC 6442542. PMID 30877254.

"Pyrgus". GBIF. Retrieved 2021-09-30.
Pyrgus page from [1]
Yellow-banded Skipper page
Pyrgus ruralis page
Edinburg WBC NE Mexico Trip
Safflower Skipper
Moths and butterflies of Europe and North Africa
English butterfly names for North America
List of Hesperiidae photographed by O. Kosterin

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