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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Infraordo: Cucujiformia
Superfamilia: Tenebrionoidea

Familia: Tenebrionidae
Subfamilia: Pimeliinae
Tribus: Scotobiini
Genera: DiastoleusEmmallodera – Pumiliofossorum – Scotobius

Flores, G.E. 2009: Revision of some types of the South American tribes Nycteliini, Praocini, and Scotobiini (Coleoptera: Tenebrionidae), with new synonymies. Zootaxa, 1985: 21–33. Abstract & excerpt
Silvestro, V.A., Giraldo-Mendoza, E.G. & Flores, G.E. 2015. Pumiliofossorum: A new genus of Scotobiini (Coleoptera: Tenebrionidae) with two new species from Peru, and a revised key for the genera of the tribe. Zootaxa 3986(4): 461–471. DOI: 10.11646/zootaxa.3986.4.5. Preview (PDF) Reference page.

Scotobiini is a tribe of darkling beetles in the family Tenebrionidae. There are about six genera in Scotobiini, found in the Neotropics.[1][2]

These genera belong to the tribe Scotobiini:

Ammophorus Guérin-Méneville, 1831
Diastoleus Solier, 1838
Emmallodera Blanchard, 1842
Leptynoderes Solier, 1838
Pumiliofossorum Silvestro & Giraldo-Mendoza, 2015
Scotobius Germar, 1823


Bouchard, Patrice; Bousquet, Yves; Aalbu, Rolf L.; Alonso-Zarazaga, Miguel A.; et al. (2021). "Review of genus-group names in the family Tenebrionidae (Insecta, Coleoptera)". ZooKeys (1050): 1–633. doi:10.3897/zookeys.1050.64217. hdl:10261/250214. PMC 8328949. PMID 34385881.

Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011). "Family-group names in Coleoptera (Insecta)". ZooKeys (88): 1–972. doi:10.3897/zookeys.88.807. PMC 3088472. PMID 21594053.

Further reading

LeConte, J. L. (1861). Classification of the Coleoptera of North America. Smithsonian Miscellaneous Collections. 3. Smithsonian Institution. doi:10.5962/bhl.title.38459. ISBN 978-0665100550.
Somerby, Ronald E.; Thomas, Donald B.; Triplehorn, Charles A. (2002). "Family 106: Tenebrionidae". In Arnett, Ross H. Jr.; Thomas, Michael C.; Skelley, Paul E.; Frank, J. Howard (eds.). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press. pp. 463–509. ISBN 978-0-8493-0954-0.

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