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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Hesperiidae
Subfamilia: Pyrginae
Tribus: Carcharodini
Genus: Staphylus
Species: S. ascalaphus – S. ascalon – S. astra – S. azteca – S. balsa – S. buena – S. caribbea – S. cartagoa – S. ceos – S. chlora – S. chlorocephala – S. coecatus – S. cordillerae – S. corumba – S. eryx – S. esmeraldus – S. evemerus – S. hayhurstii – S. huigra – S. iguala – S. imperspicua – S. incanus – S. insignis – S. kayei – S. lenis – S. lizeri – S. mazans – S. melaina – S. melangon – S. melius – S. menuda – S. minor – S. mossi – S. musculus – S. oeta – S. parvus – S. perforata – S. perna – S. punctiseparatus – S. putumayo – S. sambo – S. saxos – S. shola – S. tepeca – S. tierra – S. tingo – S. tridentis – S. tucumanus – S. tyro – S. unicornis – S. veytius – S. vincula – S. vulgata – S. zuritus
Name

Staphylus Godman & Salvin, [1896].

Type species: Helias ascalaphus Staudinger, 1875ː 118.
Synonymy

Staphylus Godman & Salvin, [1896]: 429; Evans, [1953]ː83; Hemming, 1967ː 417.
Scantilla Godman & Salvin, [1896]: 437. Synonymy in Lamas, 2004: 55.
Staphilus Spitz, 1932. Mis-spelling per Lamas, 2004: 55.
Staphylis Worth, 1998. Mis-spelling per Lamas, 2004ː 55.

References

Evans, W.H. 1951–55. A Catalogue of the American Hesperiidae in the British Museum. Pt.1 (1951): x+92pp, pls. 1–9; Pt.2 (1952): v.+178pp, pls. 10–25; Pt.3 (1953): v+246pp, pls. 26–53; Pt.4 (1955): v+499pp, pls. 54–88. Reference page.
Freeman, H.A. 1969. Systematic review of the Megathymidae. Journal of the Lepidopterists' Society 23(suppl. 1): 1–62. Reference page.
Godman, F. & Salvin, O. 1879-1901. Biologia Centrali-Americana (Lepidoptera; Rhopalocera), 3 vols. London, Taylor & Francis. Vol. I, Vol. II, Vol III (plates). Reference page.
Hemming, A. F., 1967. The generic names of the butterflies and their type species (Lepidoptera: Rhopalocera). Bulletin of the British Museum (Natural History) 1967, Suppl. 9: 1-509.Reference page.
Lamas, G. 2004. (ed.) Checklist: Part 4A. Hesperioidea - Papilionoidea. In Heppner, J.B. (ed.) Atlas of Neotropical Lepidoptera. Vol.5A, Pt.4A. Assn. for Tropical Lepidoptera/Scientific Publishers, Gainesville. 439pp. Reference page.
Miller, L.D., 1966. A New Staphylus from Costa Rica (Lepid.: Hesperiidae). Entomological News 77ː 261-264, Figs 1-3. Reference page.
Staudinger, O., 1875. Neue Lepidopteren des südamerikanischen Faunegebeits. Verh. zool.-bot. Ges. Wien. 25: 89-124. Reference page.
Steinhauser, S.R., 1989. Taxonomic notes and descriptions of new taxa in the neotropical Hesperiidae. Part I. Pyrginae. Bulletin of the Allyn Museum 127: 1–70. Reference page.

Staphylus is a genus of skippers in the family Hesperiidae found mainly in South America with some species in the Caribbean, Mexico, Central America, and southern United States.[1]
Species

Listed alphabetically.[1]

Staphylus ascalaphus (Staudinger, 1876) – mauve scallopwing
Staphylus ascalon (Staudinger, 1876)
Staphylus astra (Williams & Bell, 1940)
Staphylus azteca (Scudder, 1872) – Aztec scallopwing
Staphylus balsa (Bell, 1937)
Staphylus buena (Williams & Bell, 1940)
Staphylus caribbea (Williams & Bell, 1940)
Staphylus cartagoa (Williams & Bell, 1940)
Staphylus ceos (Edwards, 1882) – golden-headed sootywing
Staphylus chlora Evans, 1953 – green-headed sootywing
Staphylus chlorocephala (Latreille, [1824])
Staphylus coecatus (Mabille, 1891)
Staphylus cordillerae (Lindsey, 1925)
Staphylus corumba (Williams & Bell, 1940)
Staphylus eryx Evans, 1953
Staphylus esmeraldus Miller, 1966
Staphylus evemerus Godman & Salvin, [1896]
Staphylus fasciatus Hayward, 1933
Staphylus hayhurstii (Edwards, 1870) – Hayhurst's scallopwing, scalloped sootywing
Staphylus huigra (Williams & Bell, 1940)
Staphylus iguala (Williams & Bell, 1940) – Iguala sootywing
Staphylus imperspicua (Hayward, 1940)
Staphylus incanus Bell, 1932
Staphylus incisus (Mabille, 1878)
Staphylus insignis Mielke, 1980
Staphylus kayei Cock, 1996
Staphylus lenis Steinhauser, 1989
Staphylus lizeri (Hayward, 1938)
Staphylus mazans (Reakirt, [1867]) – Mazans scallopwing, southern scalloped sootywing
Staphylus melaina (Hayward, 1947)
Staphylus melangon (Mabille, 1883)
Staphylus melius Steinhauser, 1989
Staphylus menuda (Weeks, 1902)
Staphylus minor Schaus, 1902 – speckled sootywing
Staphylus mossi Evans, 1953
Staphylus musculus (Burmeister, 1875)
Staphylus oeta (Plötz, 1884) – Plötz's sootywing
Staphylus parvus Steinhauser & Austin, 1993
Staphylus perforata (Möschler, 1879)
Staphylus perna Evans, 1953
Staphylus punctiseparatus Hayward, 1933
Staphylus putumayo (Bell, 1937)
Staphylus sambo Evans, 1953
Staphylus saxos Evans, 1953
Staphylus shola Evans, 1953
Staphylus tepeca (Bell, 1942) – checkered scallopwing
Staphylus tierra Evans, 1953 – west-Mexican scallopwing
Staphylus tingo Steinhauser, 1989
Staphylus tridentis Steinhauser, 1989
Staphylus tucumanus (Plötz, 1884)
Staphylus tyro (Mabille, 1878)
Staphylus unicornis Steinhauser & Austin, 1993
Staphylus veytius Freeman, 1969 – Chiapas scallopwing
Staphylus vincula (Plötz, 1886) – mountain scallopwing
Staphylus vulgata (Möschler, 1879) – golden-snouted scallopwing

References

Staphylus, funet.fi

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