Trachypachidae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: Adephaga

Familia: Trachypachidae
Subfamiliae (3): Systolosomatinae - Trachypachinae - †Eodromeinae
Overview of genera (2)

Systolosoma – Trachypachus
+ fossil genera
Name

Trachypachidae Thomson, 1857

References

Arndt, E.; Beutel, R.G. 1995: Larval morphology of Systolosoma Solier and Trachypachus Motschulsky (Coleoptera: Trachypachidae) with phylogenetic considerations. Entomologica scandinavica, 26: 439–446. DOI: 10.1163/187631295X00099
Kavanaugh, D.H.; Erwin, T.L.; Maddison, D.R. (in prep): The Trachypachini (Coleoptera: Carabidae): taxonomy and phylogeny, based on a study of adults and larvae.
Lorenz, W. 2005: A systematic list of extant ground beetles of the World (Coleoptera "Geadephaga": Trachypachidae and Carabidae, incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. (Tutzing) ISBN 3-933 896-06-1 first page of page by page pdfs
Martínez-Navarro, E.M.; Serrano, J.; Galián, J. 2011: Chromosomes of Trachypachus Motschulsky and the evolution of the ancestral adephagan karyotype (Coleoptera). Journal of Zoological Systematics and Evolutionary Research, 49(3): 251–255. DOI: 10.1111/j.1439-0469.2010.00613.x
Wang, B.; Zhang, H.; Ponomarenko, A.G. 2012: Mesozoic Trachypachidae (Insecta: Coleoptera) from China. Palaeontology, 55(2): 341–353. DOI: 10.1111/j.1475-4983.2012.01128.x
Atlas of the family Trachypachidae of Russia
Catalogue of Palearctic Coleoptera. Vol. 1 ed. I. Lobl, & A. Smetana, Apollo Books, Stenstrup, Denmark, 2003 and 2004
ISBN 87-88757-73-0, p.33

Links

Trachypachidae in SYNOPSIS OF THE DESCRIBED COLEOPTERA OF THE WORLD
Tree of Life Web Project. 1995. Trachypachidae. Version 01 January 1995 (temporary) in The Tree of Life Web Project
EoL

Vernacular names
English: False Ground Beetles
한국어: 어리강변먼지벌레과
norsk: Urløpebiller
русский: Трахипахиды

The Trachypachidae (sometimes known as false ground beetles) are a family of beetles that generally resemble small ground beetles, but that are distinguished by the large coxae of their rearmost legs. There are only six known extant species in the family, with four species of Trachypachus found in northern Eurasia and northern North America, and two species of Systolosoma in Chile. They were much more diverse in the past, with many members belonging to the extinct subfamily Eodromeinae,[1] the first fossils known of this family are of the genera Petrodromeus and Permunda from the Permian-Triassic boundary of Russia.[2]

Their habits are similar to those of the ground beetles; they are usually found in the leaf litter of conifer forests.

To quote G. E. Ball, "the most interesting thing about this small family is its uncertain phylogenetic relationships"; it has been classified as a subfamily of Carabidae, and as a family been placed in several different positions relative to other beetle families.
Species

Subfamily Trachypachinae
Genus Systolosoma
Systolosoma breve
Systolosoma lateritium
Genus Trachypachus
Trachypachus holmbergi
Trachypachus gibbsii
Trachypachus zetterstedi
Trachypachus slevini
Subfamily †Eodromeinae
Genus †Eodromeus
Genus †Sogdodromeus
†Sogdodromeus altus
Genus †Petrodromeus
†Petrodromeus asiaticus
†Petrodromeus minor
Genus †Permunda
†Permunda nana
Genus †Unda
Genus †Sinodromeus
†Sinodromeus liutiaogouensis
Genus †Praedodromeus
†Praedodromeus sangiorgiensis

References

Wang, Bo; Zhang, Haichun; Ponomarenko, Alexandr G. (2012). "Mesozoic Trachypachidae (Insecta: Coleoptera) from China". Palaeontology. 55 (2): 341–353. doi:10.1111/j.1475-4983.2012.01128.x. ISSN 0031-0239.

Ponomarenko, A. G.; Volkov, A. N. (2013). "Ademosynoides asiaticus Martynov, 1936, the earliest known member of an extant beetle family (Insecta, Coleoptera, Trachypachidae)". Paleontological Journal. 47 (6): 601–606. doi:10.1134/S0031030113060063. ISSN 0031-0301.

George E. Ball, "Trachypachidae", in Ross H. Arnett, Jr. and Michael C. Thomas, American Beetles (CRC Press, 2001), vol. 1
L. Strada, M. Montagna, and A. Tintori. 2014. A new genus and species of the family Trachypachidae (Coleoptera, Adephaga) from the upper Ladinian (Middle Triassic) of Monte San Giorgio. Rivista Italiana di Paleontologia e Stratigrafia 120:183-190