Trigonopterus

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Ordo: Coleoptera
Subordo: Polyphaga
Superfamilia: Curculionoidea

Familia: Curculionidae
Subfamilia: Molytinae
Tribus: Cryptorhynchini
Subtribus: Tylodina
Genus: Trigonopterus
Species (472):
A

T. abnormis – T. acuminatus – T. acutus – T. adspersus – T. aeneipennis – T. aeneomicans – T. aeneoniveus – T. aeneus – T. aequalis – T. agathis – T. agilis – T. alaspurwensis – T. albidosparsus – T. albopunctatus – T. allaetus – T. allopatricus – T. allotopus – T. ambangensis – T. ampanensis – T. amphoralis – T. ampliatus – T. amplipennis – T. analis – T. ancora – T. ancoruncus – T. angulatus – T. angulicollis – T. angustus – T. anthracinus – T. anthrax – T. apicalis – T. arachnobas – T. arcanus – T. argopurensis – T. arjunensis – T. armatus – T. armipes – T. artemis – T. ascendens – T. asper – T. asterix – T. athertonensis – T. attenboroughi – T. atuf – T. augur – T. australinasutus – T. australis –
B

T. bakeri – T. baliensis – T. balimensis – T. barbipes – T. basalis – T. basimaculatus – T. batukarensis – T. bawangensis – T. bicolor – T. biguttatoides – T. biguttatus – T. binodulus – T. binotatus – T. bisignatus – T. bisinuatus – T. bonthainensis – T. boolbunensis – T. bornensis – T. bryani –
C

T. caesipes – T. cahyoi – T. carinatus – T. carinirostris – T. castaneipennis – T. caudatus – T. celebensis – T. chewbacca – T. cirripes – T. collaris – T. conformis – T. conicus – T. constrictus – T. convexus – T. cooktownensis – T. coriarius – T. corona – T. costatulus – T. costatus – T. costicollis – T. costipennis – T. crassicornis – T. crenulatus – T. cribratus – T. cribricollis – T. cricki – T. crinipes – T. cuneatus – T. cuneipennis – T. cuprescens – T. cupreus – T. curtus – T. curvipes – T. cyclopensis –
D

T. dacrycarpi – T. daintreensis – T. dakoensis – T. darwini – T. daun – T. dehiscens – T. delapan – T. densatus – T. dentipes – T. dentirostris – T. deplanatus – T. diengensis – T. difficilis – T. difformis – T. dilaticollis – T. dimorphus – T. discoidalis – T. disruptus – T. diversicollis – T. dromedarius – T. dua – T. duabelas – T. durus –
E

T. ebriosus – T. echinatus – T. echinus – T. edaphus – T. ejaculatorius – T. ellipticus – T. empat – T. enam – T. ephippiatus – T. eremitus – T. euops – T. evanidus – T. ewok –
F

T. femoralis – T. femoridens – T. ferrugineus – T. finniganensis – T. fissitarsis – T. flavomaculatus – T. florensis – T. foveatus – T. fraterculus – T. fulgidus – T. fulvicornis – T. fuscipes – T. fusiformis –
G

T. garradungensis – T. gedensis – T. gibbirostris – T. glaber – T. globatus – T. gonatoceros – T. gracilipes – T. grandicollis – T. granum – T. gundala –
H

T. halimunensis – T. hartleyensis – T. hasenpuschi – T. hassleri – T. heberti – T. helios – T. heteropunctatus – T. hirsutus – T. hitoloorum – T. honestus – T. honjensis – T. hoppla – T. humeralis – T. humilis – T. hypocrita –
I

T. idefix – T. ijensis – T. illex – T. illitus – T. imitatus – T. impar – T. impressicollis – T. incendium – T. inclusus – T. incognitus – T. indigenus – T. inflatus – T. inhonestus – T. insignis – T. insignoides – T. insularis – T. interpositus – T. invalidus – T. irregularis – T. ixodiformis –
J

T. jasminae – T. javensis – T. jekelii –
K

T. kakimerah – T. kalimantanensis – T. kanawiorum – T. katopasensis – T. katayoi – T. kintamanensis – T. klabatensis – T. klatakanensis – T. kolakensis – T. kotamobagensis – T. koveorum – T. kumbang – T. kurandensis – T. kurulu – T. kuscheli –
L

T. laetus – T. laevigatus – T. lampros – T. lampungensis – T. laratensis – T. latipennis – T. latipes – T. lekiorum – T. lescheni – T. lewisensis – T. lima – T. lineatus – T. lineellus – T. lombokensis – T. lompobattangensis – T. luwukensis –
M

T. maculatus – T. mahawuensis – T. manadensis – T. mangkutanensis – T. matakensis – T. matalibaruensis – T. melas – T. merophysioides – T. merubetirensis – T. mesai – T. mesehensis – T. micans – T. micros – T. mimicus – T. minahassae – T. minutus – T. misellus – T. moatensis – T. modoindingensis – T. moduai – T. mons – T. montanus – T. monteithi – T. monticola – T. montivagus – T. moreaorum – T. morokensis – T. mossmanensis – T. myops –
N

T. nangiorum – T. nanus – T. nasutus – T. neglectus – T. nitidulus – T. nothofagorum –
O

T. obelix – T. oberprieleri – T. oblitus – T. oblongus – T. obnixus – T. obsidianus – T. obsoletus – T. ocladiiformis – T. ovalipunctatus – T. ovatulus – T. ovatus – T. oviformis –
P

T. pagaranganensis – T. palawanensis – T. palopensis – T. pangandaranensis – T. paracollaris – T. paraflorensis – T. paramoduai – T. pararugosus – T. parasumbawensis – T. parumsquamosus – T. parvulus – T. parvus – T. paucisquamosus – T. pauper – T. pauxillus – T. payungensis – T. pembertoni – T. pendolensis – T. perpolitus – T. phoenix – T. pilosipes – T. plicicollis – T. politoides – T. politus – T. pomberimbensis – T. pompangeensis – T. porcatus – T. porg – T. posoensis – T. prismae – T. procurtus – T. proximus – T. pseudallotopus – T. pseudanalis – T. pseudinsignis – T. pseudoflorensis – T. pseudofulvicornis – T. pseudogranum – T. pseudomanadensis – T. pseudonasutus – T. pseudosimulans – T. pseudosumbawensis – T. pseudovatulus – T. pseudovalipunctatus – T. ptolycoides – T. pulchellus – T. pulicaris – T. pullus – T. pumilus – T. punctatoseriatus – T. puncticollis – T. punctulatus – T. pusillus – T. puspoi –
Q

T. quadrimaculatus –
R

T. ragaorum – T. ranakensis – T. rantepao – T. relictus – T. reticulatus – T. rhinoceros – T. rhombiformis – T. rhomboidalis – T. rinjaniensis – T. robertsi – T. roensis – T. rosichoni – T. rostralis – T. rotundatus – T. rotundulus – T. rubidus – T. rubiginosus – T. rubripennis – T. rudis – T. rufibasis – T. rufipennis – T. rufipes – T. rufipterus – T. rufithorax – T. rugosostriatus – T. rugosus – T. rutengensis –
S

T. saltator – T. salubris – T. samoanus – T. sampunensis – T. sampuragensis – T. santubongensis – T. sarinoi – T. sasak – T. satu – T. satyrus – T. scabripes – T. scabrosus – T. scaphiformis – T. scaphioides – T. scharfi – T. schulzi – T. scissops – T. scitulus – T. sculptirostris – T. sculpturatus – T. sebelas – T. seclusus – T. sejunctus – T. selaruensis – T. selayarensis – T. sellatus – T. sembilan – T. semicribrosus – T. semirubrus – T. sepuluh – T. seriatus – T. serratifemur – T. serratipes – T. serripes – T. seticnemis – T. setifer – T. setipoides – T. setipus – T. sexmaculatus – T. signicollis – T. silaliensis – T. silvestris – T. silvicola – T. similis – T. simulans – T. singkawangensis – T. singularis – T. sinuatus – T. soiorum – T. solidus – T. sordidus – T. squalidulus – T. squalidus – T. squamirostris – T. squamosus – T. striatipennis – T. striatus – T. strigatus – T. strombosceroides – T. subglabratus – T. submetallicus – T. subrubricollis – T. sulawesiensis – T. sulcatus – T. sumatrensis – T. sumbawensis – T. sundaicus – T. sutrisnoi – T. suturaelevata – T. suturalis – T. suturatus – T. syarbis –
T

T. taenzleri – T. talpa – T. tanah – T. tanimbarensis – T. tatorensis – T. taurekaorum – T. tejokusumoi – T. telagensis – T. tenuipes – T. tepalensis – T. terraereginae – T. tialeorum – T. tibialis – T. tiga – T. toboliensis – T. tolitoliensis – T. tomohonensis – T. toraja – T. tounaensis – T. tridentatus – T. trigonopterus – T. triradiatus – T. tujuh –
U

T. ujungkulonensis – T. unicolor – T. uniformis – T. unyil –
V

T. vandekampi – T. vanus – T. variabilis – T. variolosus – T. velaris – T. verrucosus – T. vicinus – T. viduus – T. violaceus – T. viridescens – T. volcanorum – T. vossi – T. vulcanicus –
W

T. wallacei – T. wamenaensis – T. wangiwangiensis – T. wariorum – T. watsoni – T. wolffi –
Y

T. yoda – T. yorkensis –
Z

T. zonatus – T. zygops
Name

Trigonopterus Fauvel, 1862: 157

Type species: Trigonopterus insignis Fauvel, 1862, by monotypy.
Synonyms

Trigonus Fauvel, 1868 [new replacement name for Trigonopterus Fauvel, 1862]
Idotasia Pascoe, 1871
Type species: Idotasia nasuta Pascoe, 1871
Fixation: subsequent designation by Riedel (2011: 2)
Eurysia Pascoe, 1885
Microgymnapterus Voss, 1960
Authority for current placement: Riedel, 2011: 3 ('Syn.n.')
Mimidotasia Voss, 1960
Authority for current placement: Riedel, 2011: 3 ('Syn.n.')

Trigonopterus vandekampi from New Guinea

References

Fauvel, A. 1862. Coléoptères de la Nouvelle-Calédonie, recueillis par M. E. Déplanche, chirurgien de la marine impériale (1858-59-60). Bulletin de la Société Linnéenne de Normandie 7: 120–185. BHL Reference page. [original description: p. 157]
Fauvel, A. 1868. Catalogue des Coléoptères de la Nouvelle Calédonie et dépendances avec descriptions, notes et synonymies nouvelles. Bulletin de la Société Linnéenne de Normandie (2) 1: 172–209. BHL Reference page. [see p. 196]
Narakusumo, R.P., Balke, M. & Riedel, A. 2019. Seven new species of Trigonopterus Fauvel (Coleoptera, Curculionidae) from the Tanimbar Archipelago. ZooKeys 888: 75-93. DOI: 10.3897/zookeys.888.38642. Reference page.
Narakusumo, R.P. & Riedel, A. 2021. Twenty-eight new species of Trigonopterus Fauvel (Coleoptera, Curculionidae) from Central Sulawesi. Zookeys 1065ː 29–79. DOI: 10.3897/zookeys.1065.71680 Open access Reference page.
Rheinheimer, J. 2013. Trigonopterus Fauvel und eine neue Cryptorhynchinen-Gattung aus Neukaledonien (Coleoptera: Curculionidae<: Cryptorhynchinae). Koleopterologische rundschau 83: 195–255. [Not seen] Reference page.
Riedel, A. 2010. One of a thousand - a new species of Trigonopterus (Coleoptera, Curculionidae, Cryptorhynchinae) from New Guinea. Zootaxa 2403: 59–68. Preview
Riedel, A. 2011. The weevil genus Trigonopterus Fauvel (Coleoptera, Curculionidae) and its synonyms—a taxonomic study on the species tied to its genus-group names. Zootaxa 2977: 1–49. Preview
Riedel, A., Daawia, D. & Balke, M. 2010. Deep cox1 divergence and hyperdiversity of Trigonopterus weevils in a New Guinea mountain range (Coleoptera, Curculionidae). Zoologica scripta 39(1): 63–74. DOI: 10.1111/j.1463-6409.2009.00404.x
Riedel, A. & Narakusumo, R.P. 2019. One hundred and three new species of Trigonopterus weevils from Sulawesi. Zookeys, 828: 1–153. DOI: 10.3897/zookeys.828.32200 Reference page.
Riedel, A., Sagata, K., Surbakti, S., Tänzler, R. & Balke, M. 2013. One hundred and one new species of Trigonopterus weevils from New Guinea. ZooKeys 280: 1–150. DOI: 10.3897/zookeys.280.3906 Reference page.
Riedel, A., Tänzler, R., Balke, M., Rahmadi, C. & Suhardjono, Y.R. 2014. Ninety-eight new species of Trigonopterus weevils from Sundaland and the Lesser Sunda Islands. ZooKeys 467: 1–162. DOI: 10.3897/zookeys.467.8206 Reference page.
Riedel, A. & Tänzler, R. 2016. Revision of the Australian species of the weevil genus Trigonopterus Fauvel. ZooKeys 556: 97–162. DOI: 10.3897/zookeys.556.6126. Reference page.
Van Dam, M.H., Laufa, R. & Riedel, A. 2016. Four new species of Trigonopterus Fauvel from the island of New Britain (Coleoptera, Curculionidae). ZooKeys 582: 129-141. DOI: 10.3897/zookeys.582.7709. Reference page.

Trigonopterus is a genus of flightless weevils placed in the Cryptorhynchinae of Curculionidae. It is distributed in Australia, Indonesia and Melanesia. About 90 species had been formally described until March 2013, when a single paper more than doubled this number,[2] agreeing with previous studies[3] and a systematic barcoding study[4] that many more species have yet to be described. As of October 2021, there were 441 described species.

The center of its diversity appears to be New Guinea where 51 or more species can be found in a single locality.[5] Many of them are very similar to each other, but male genital characters and DNA barcoding allow a safe identification.

In January 2016, a paper was published that revised the Australian species within this genus. That paper described 24 new species and indicated the potential for more research into undescribed Australian species within this genus.[6][7]

In April 2016, a paper was published that described four new species in this genus from the island of New Britain. The paper postulated that more species of this genus presently unknown to science may exist on the island. It emphasised the importance of prioritising further research due to the ongoing destruction of the habitat of these species.[8]

Further papers described 133 new species from Sulawesi, where only one species (T. fulvicornis) had previously been recorded.[9][10]

The genus can be diagnosed among wingless cryptorhynchine weevils by the absence of a metanepisternum and by a synapomorphic structure of the tarsus with minute claws and a deeply incavated articulation of tarsomere 4. The metathoracic spiracle located externally at the side of the metaventrite is a unique feature and may ensure sufficient respiration during thanatosis.[11]

Trigonopterus species inhabit primary tropical forests, both on foliage and edaphic in the litter layer. They have a marked tendency to endemism with many species only known from a single locality. Their primary defence against predators is apparent death or thanatosis. An animated 3D model of a Trigonopterus weevil reveals a number of mechanisms to maintain a stable defensive position.[12]

Biological screw joint
Main article: Biological screw joint
100 Trigonopterus species described in 2013

The arthropod hip-leg joint consists of two parts - the coxa (or the hip) and the trochanter (or the head of the arthropod leg femur).[13] The coxa, in the case of Trigonopterus oblongus, resembles a nut, and it has a thread running along its inner surface with an angular span of 345°.[14][13] The trochanter resembles the screw.[13] It is rod-shaped with a large external spiral flange, having an angular span of 410°, in excess of a full circle, which functions as a thread.[14] When the leg muscles of a beetle are stretched, the screw turns. Though the screw-thread provide for very large angular rotation, the front legs are capable of rotating by 90°, while their hind legs can rotate by 130°.[15]
Evolution

The screw-and-nut system has now been found to be present in all 15 weevil species examined by the scientists and appears to be a hitherto unknown anatomical feature of weevils.[16][15] It has been estimated that weevils evolved this system about 100 million years ago. It is surmised that the development of this feature provided additional flexibility which permitted weevils to improve their climbing abilities, helped them keep steady when at rest, and to give a stronger leverage for piercing by the snout.[16]
Species

Trigonopterus abnormis Riedel, 2019
Trigonopterus acuminatus Riedel, 2014
Trigonopterus acutus Narakusumo & Riedel, 2021
Trigonopterus adspersus Riedel, 2019
Trigonopterus aeneipennis Riedel, 2013
Trigonopterus aeneomicans Riedel, 2014
Trigonopterus aeneoniveus Fairmaire, 1879
Trigonopterus aeneus Riedel, 2013
Trigonopterus aequalis (Pascoe, 1872)
Trigonopterus agathis Riedel, 2013
Trigonopterus agilis Riedel, 2013
Trigonopterus alaspurwensis Riedel, 2014
Trigonopterus albidosparsus (Lea, 1913)
Trigonopterus albopunctatus Rheinheimer, 2013
Trigonopterus allaetus Riedel, 2016
Trigonopterus allopatricus Riedel, 2014
Trigonopterus allotopus Riedel, 2014
Trigonopterus ambangensis Riedel, 2019
Trigonopterus ampanensis Riedel, 2019
Trigonopterus amphoralis Marshall, 1925
Trigonopterus ampliatus (Pascoe, 1885)
Trigonopterus amplipennis Riedel, 2013
Trigonopterus analis Riedel, 2019
Trigonopterus ancora Narakusumo & Riedel, 2021
Trigonopterus ancoruncus Riedel, 2013
Trigonopterus angulatus Riedel, 2013
Trigonopterus angulicollis Riedel, 2014
Trigonopterus angustus Riedel, 2013
Trigonopterus anthracinus (Voss, 1960)
Trigonopterus anthrax Fairmaire, 1881
Trigonopterus apicalis Riedel, 2013
Trigonopterus arachnobas Riedel, 2019
Trigonopterus arcanus Narakusumo & Riedel, 2021
Trigonopterus argopurensis Riedel, 2014
Trigonopterus arjunensis Riedel, 2014
Trigonopterus armatus Riedel, 2013
Trigonopterus armipes Riedel, 2019
Trigonopterus artemis Riedel, 2019
Trigonopterus ascendens Riedel, 2013
Trigonopterus asper Riedel, 2014
Trigonopterus asterix Riedel, 2019
Trigonopterus athertonensis Riedel, 2016
Trigonopterus attenboroughi Riedel, 2014
Trigonopterus atuf Narakusumo & Riedel, 2019
Trigonopterus augur Riedel, 2013
Trigonopterus australinasutus Riedel, 2016
Trigonopterus australis Riedel, 2016
Trigonopterus bakeri (Hustache, 1925)
Trigonopterus baliensis Riedel, 2014
Trigonopterus balimensis Riedel, 2013
Trigonopterus barbipes Riedel, 2019
Trigonopterus basalis Riedel, 2013
Trigonopterus basimaculatus (Voss, 1960)
Trigonopterus batukarensis Riedel, 2014
Trigonopterus bawangensis Riedel, 2014
Trigonopterus bicolor Marshall, 1921
Trigonopterus biguttatoides Rheinheimer, 2013
Trigonopterus biguttatus Rheinheimer, 2013
Trigonopterus binodulus Riedel, 2014
Trigonopterus binotatus Marshall, 1921
Trigonopterus bisignatus Riedel, 2016
Trigonopterus bisinuatus Riedel, 2016
Trigonopterus bonthainensis Riedel, 2019
Trigonopterus boolbunensis Riedel, 2016
Trigonopterus bornensis Riedel, 2014
Trigonopterus bryani Marshall, 1931
Trigonopterus caesipes Marshall, 1931
Trigonopterus cahyoi Riedel, 2014
Trigonopterus carinatus Rheinheimer, 2013
Trigonopterus carinirostris Riedel, 2019
Trigonopterus carolineae Zimmerman, 1941
Trigonopterus castaneipennis Riedel, 2019
Trigonopterus caudatus Heller, 1920
Trigonopterus celebensis Riedel, 2019
Trigonopterus chewbacca Van Dam & Riedel, 2016
Trigonopterus cirripes Riedel, 2019
Trigonopterus collaris Riedel, 2019
Trigonopterus conformis Riedel, 2013
Trigonopterus conicus Rheinheimer, 2013
Trigonopterus constrictus Riedel, 2013
Trigonopterus convexus (Faust, 1898)
Trigonopterus cooktownensis Riedel, 2016
Trigonopterus coriarius Heller, 1920
Trigonopterus corona Narakusumo & Riedel, 2021
Trigonopterus costatulus Riedel, 2019
Trigonopterus costatus Riedel, 2013
Trigonopterus costicollis Riedel, 2013
Trigonopterus costipennis Riedel, 2014
Trigonopterus crassicornis Riedel, 2013
Trigonopterus crenulatus Riedel, 2019
Trigonopterus cribratus (Faust, 1899)
Trigonopterus cribrellicollis Fairmaire, 1881
Trigonopterus cricki Riedel, 2019
Trigonopterus crinipes Marshall, 1931
Trigonopterus cuneatus (Faust, 1899)
Trigonopterus cuneipennis Riedel, 2013
Trigonopterus cuprescens Riedel, 2014
Trigonopterus cupreus Riedel, 2014
Trigonopterus curtus (Voss, 1960)
Trigonopterus curvipes Riedel, 2019
Trigonopterus cyclopensis Riedel, 2013
Trigonopterus dacrycarpi Riedel, 2014
Trigonopterus daintreensis Riedel, 2016
Trigonopterus dakoensis Narakusumo & Riedel, 2021
Trigonopterus darwini Riedel, 2019
Trigonopterus daun Narakusumo & Riedel, 2021
Trigonopterus delapan Riedel, 2014
Trigonopterus densatus (Faust, 1899)
Trigonopterus dentipes Riedel, 2014
Trigonopterus dentirostris Riedel, 2013
Trigonopterus deplanatus Riedel, 2016
Trigonopterus diengensis Riedel, 2014
Trigonopterus difficilis (Faust, 1899)
Trigonopterus difformis Rheinheimer, 2013
Trigonopterus dilaticollis (Faust, 1898)
Trigonopterus dimorphus Riedel, 2014
Trigonopterus discoidalis Riedel, 2013
Trigonopterus disruptus Riedel, 2014
Trigonopterus diversicollis Heller, 1920
Trigonopterus dromedarius Riedel, 2013
Trigonopterus dua Riedel, 2014
Trigonopterus duabelas Riedel, 2014
Trigonopterus durus Riedel, 2013
Trigonopterus ebriosus (Pascoe, 1871)
Trigonopterus echinatus Riedel, 2014
Trigonopterus echinus Riedel, 2013
Trigonopterus edaphus Riedel, 2013
Trigonopterus ejaculatorius Riedel, 2019
Trigonopterus ellipticus (Pascoe, 1871)
Trigonopterus empat Riedel, 2014
Trigonopterus enam Riedel, 2014
Trigonopterus ephippiatus (Faust, 1899)
Trigonopterus eremitus Riedel, 2013
Trigonopterus euops Riedel, 2013
Trigonopterus evanidus Rheinheimer, 2013
Trigonopterus ewok Narakusumo & Riedel, 2021
Trigonopterus femoralis (Faust, 1898)
Trigonopterus femoridens Rheinheimer, 2013
Trigonopterus ferrugineus Riedel, 2013
Trigonopterus finniganensis Riedel, 2016
Trigonopterus fissitarsis Riedel, 2014
Trigonopterus flavomaculatus (Voss, 1960)
Trigonopterus florensis Riedel, 2014
Trigonopterus foveatus Riedel, 2014
Trigonopterus fraterculus Riedel, 2016
Trigonopterus fulgidus Riedel, 2014
Trigonopterus fulvicornis (Pascoe, 1885)
Trigonopterus fuscipes Riedel, 2019
Trigonopterus fusiformis Riedel, 2013
Trigonopterus garradungensis Riedel, 2016
Trigonopterus gedensis Riedel, 2014
Trigonopterus gibbirostris (Faust, 1899)
Trigonopterus glaber Riedel, 2013
Trigonopterus globatus (Voss, 1960)
Trigonopterus gonatoceros Riedel, 2013
Trigonopterus gracilipes Riedel, 2019
Trigonopterus granum Riedel, 2013
Trigonopterus gundala Narakusumo & Riedel, 2021
Trigonopterus halimunensis Riedel, 2014
Trigonopterus hartleyensis Riedel, 2016
Trigonopterus hasenpuschi Riedel, 2016
Trigonopterus hassleri Rheinheimer, 2013
Trigonopterus heberti Riedel, 2019
Trigonopterus helios Riedel, 2013
Trigonopterus heteropunctatus Heller, 1920
Trigonopterus hirsutus Riedel, 2019
Trigonopterus hitoloorum Riedel, 2013
Trigonopterus honjensis Riedel, 2014
Trigonopterus hoppla Narakusumo & Riedel, 2021
Trigonopterus humilis Riedel, 2019
Trigonopterus hypocrita Riedel, 2019
Trigonopterus idefix Riedel, 2019
Trigonopterus ijensis Riedel, 2014
Trigonopterus illex (Faust, 1899)
Trigonopterus illitus (Faust, 1899)
Trigonopterus imitatus Riedel, 2013
Trigonopterus impar (Faust, 1898)
Trigonopterus impressicollis Riedel, 2019
Trigonopterus incendium Riedel, 2019
Trigonopterus inclusus (Pascoe, 1871)
Trigonopterus incognitus Riedel, 2019
Trigonopterus indigenus Riedel, 2019
Trigonopterus inflatus Riedel, 2013
Trigonopterus inhonestus Riedel, 2019
Trigonopterus insignis Fauvel, 1862
Trigonopterus insularis Riedel, 2013
Trigonopterus interpositus (Voss, 1960)
Trigonopterus invalidus Riedel, 2019
Trigonopterus irregularis Riedel, 2013
Trigonopterus ixodiformis Riedel, 2013
Trigonopterus jasminae Riedel, 2019
Trigonopterus javensis Riedel, 2014
Trigonopterus jekelii (Fauvel, 1867)
Trigonopterus kakimerah Narakusumo & Riedel, 2021
Trigonopterus kalimantanensis Riedel, 2014
Trigonopterus kanawiorum Riedel, 2013
Trigonopterus katayoi Riedel, 2013
Trigonopterus katopasensis Narakusumo & Riedel, 2021
Trigonopterus kintamanensis Riedel, 2014
Trigonopterus klabatensis Riedel, 2019
Trigonopterus klatakanensis Riedel, 2014
Trigonopterus kolakensis Riedel, 2019
Trigonopterus kotamobagensis Riedel, 2019
Trigonopterus koveorum Riedel, 2013
Trigonopterus kumbang Narakusumo & Riedel, 2019
Trigonopterus kurandensis Riedel, 2016
Trigonopterus kurulu Riedel, 2013
Trigonopterus kuscheli Rheinheimer, 2013
Trigonopterus laetus (Lea, 1913)
Trigonopterus laevigatus Riedel, 2019
Trigonopterus lampros Riedel, 2019
Trigonopterus lampungensis Riedel, 2014
Trigonopterus laratensis Narakusumo & Riedel, 2019
Trigonopterus latipennis Riedel, 2019
Trigonopterus latipes Riedel, 2014
Trigonopterus lekiorum Riedel, 2013
Trigonopterus lescheni Rheinheimer, 2013
Trigonopterus lewisensis Riedel, 2016
Trigonopterus lima Riedel, 2014
Trigonopterus lineatus Riedel, 2013
Trigonopterus lineellus Riedel, 2013
Trigonopterus lombokensis Riedel, 2014
Trigonopterus lompobattangensis Riedel, 2019
Trigonopterus luwukensis Riedel, 2019
Trigonopterus maculatus Riedel, 2013
Trigonopterus mahawuensis Riedel, 2019
Trigonopterus manadensis Riedel, 2019
Trigonopterus mangkutanensis Riedel, 2019
Trigonopterus matakensis Narakusumo & Riedel, 2021
Trigonopterus matalibaruensis Riedel, 2019
Trigonopterus melas (Faust, 1899)
Trigonopterus mendax Marshall, 1931
Trigonopterus merophysioides Fairmaire, 1881
Trigonopterus merubetirensis Riedel, 2014
Trigonopterus mesai Riedel, 2019
Trigonopterus mesehensis Riedel, 2014
Trigonopterus micans Riedel, 2014
Trigonopterus mimicus Riedel, 2013
Trigonopterus minahassae Riedel, 2019
Trigonopterus minutus (Voss, 1960)
Trigonopterus misellus Riedel, 2014
Trigonopterus moatensis Riedel, 2019
Trigonopterus modoindingensis Riedel, 2019
Trigonopterus moduai Narakusumo & Riedel, 2021
Trigonopterus mons Narakusumo & Riedel, 2021
Trigonopterus montanus Riedel, 2016
Trigonopterus monteithi Riedel, 2016
Trigonopterus monticola Riedel, 2013
Trigonopterus montivagus Riedel, 2013
Trigonopterus moreaorum Riedel, 2013
Trigonopterus morokensis (Faust, 1899)
Trigonopterus mossmanensis Riedel, 2016
Trigonopterus myops Riedel, 2013
Trigonopterus nangiorum Riedel, 2013
Trigonopterus nanus Riedel, 2019
Trigonopterus nasutus (Pascoe, 1871)
Trigonopterus neglectus (Faust, 1899)
Trigonopterus nitidulus Riedel, 2019
Trigonopterus nothofagorum Riedel, 2013
Trigonopterus obelix Riedel, 2019
Trigonopterus oberprieleri Riedel, 2016
Trigonopterus oblongus (Pascoe, 1885)
Trigonopterus obnixus (Faust, 1899)
Trigonopterus obsidianus Van Dam & Riedel, 2016
Trigonopterus ovalipunctatus Riedel, 2019
Trigonopterus ovatulus Riedel, 2019
Trigonopterus ovatus Riedel, 2013
Trigonopterus oviformis Riedel, 2013
Trigonopterus pagaranganensis Riedel, 2019
Trigonopterus palawanensis Riedel, 2014
Trigonopterus palopensis Riedel, 2019
Trigonopterus pangandaranensis Riedel, 2014
Trigonopterus paracollaris Riedel, 2019
Trigonopterus paraflorensis Riedel, 2014
Trigonopterus paramoduai Narakusumo & Riedel, 2021
Trigonopterus pararugosus Riedel, 2014
Trigonopterus parasumbawensis Riedel, 2014
Trigonopterus parumsquamosus Riedel, 2013
Trigonopterus parvulus Riedel, 2013
Trigonopterus parvus Rheinheimer, 2013
Trigonopterus paucisquamosus (Heller, 1915)
Trigonopterus pauper Riedel, 2019
Trigonopterus pauxillus Riedel, 2014
Trigonopterus payungensis Riedel, 2014
Trigonopterus pembertoni (Zimmerman, 1938)
Trigonopterus pendolensis Riedel, 2019
Trigonopterus phoenix Riedel, 2013
Trigonopterus pilosipes Rheinheimer, 2013
Trigonopterus plicicollis Riedel, 2013
Trigonopterus politoides Riedel, 2013
Trigonopterus politus (Faust, 1899)
Trigonopterus pomberimbensis Narakusumo & Riedel, 2021
Trigonopterus pompangeensis Narakusumo & Riedel, 2021
Trigonopterus porcatus Riedel, 2014
Trigonopterus porg Narakusumo & Riedel, 2019
Trigonopterus posoensis Riedel, 2019
Trigonopterus prismae Riedel, 2019
Trigonopterus procurtus Riedel, 2019
Trigonopterus proximus (Voss, 1960)
Trigonopterus pseudallotopus Riedel, 2019
Trigonopterus pseudanalis Riedel, 2019
Trigonopterus pseudinsignis Rheinheimer, 2013
Trigonopterus pseudoflorensis Riedel, 2014
Trigonopterus pseudogranum Riedel, 2013
Trigonopterus pseudomanadensis Riedel, 2019
Trigonopterus pseudonasutus Riedel, 2013
Trigonopterus pseudosimulans Riedel, 2019
Trigonopterus pseudosumbawensis Riedel, 2014
Trigonopterus pseudofulvicornis Riedel, 2019
Trigonopterus pseudovalipunctatus Riedel, 2019
Trigonopterus pseudovatulus Riedel, 2019
Trigonopterus ptolycoides Riedel, 2013
Trigonopterus pulchellus (Pascoe, 1885)
Trigonopterus pulicaris (Pascoe, 1885)
Trigonopterus pullus (Voss, 1960)
Trigonopterus pumilus Riedel, 2019
Trigonopterus punctatoseriatus Riedel, 2014
Trigonopterus puncticollis Van Dam & Riedel, 2016
Trigonopterus punctulatus Riedel, 2013
Trigonopterus pusillus (Faust, 1899)
Trigonopterus puspoi Narakusumo & Riedel, 2021
Trigonopterus quadrimaculatus Rheinheimer, 2013
Trigonopterus ragaorum Riedel, 2013
Trigonopterus ranakensis Riedel, 2014
Trigonopterus rantepao Riedel, 2019
Trigonopterus relictus Riedel, 2014
Trigonopterus reticulatus Riedel, 2019
Trigonopterus rhinoceros Riedel, 2013
Trigonopterus rhomboidalis Riedel, 2013
Trigonopterus rhombiformis Riedel, 2019
Trigonopterus rinjaniensis Riedel, 2014
Trigonopterus robertsi Riedel, 2016
Trigonopterus roensis Riedel, 2014
Trigonopterus rosichoni Narakusumo & Riedel, 2021
Trigonopterus rostralis (Lea, 1928)
Trigonopterus rotundatus Riedel, 2019
Trigonopterus rotundulus Riedel, 2019
Trigonopterus rubidus Narakusumo & Riedel, 2021
Trigonopterus rubiginosus Riedel, 2013
Trigonopterus rubripennis Riedel, 2013
Trigonopterus rudis Riedel, 2019
Trigonopterus rufibasis Riedel, 2013
Trigonopterus rufipennis (Pascoe, 1885)
Trigonopterus rufipes Riedel, 2019
Trigonopterus rufipterus Rheinheimer, 2013
Trigonopterus rufithorax Rheinheimer, 2013
Trigonopterus rugosostriatus Riedel, 2014
Trigonopterus rugosus Riedel, 2014
Trigonopterus rutengensis Riedel, 2014
Trigonopterus saltator Riedel, 2014
Trigonopterus salubris (Faust, 1898)
Trigonopterus sampunensis Riedel, 2019
Trigonopterus sampuragensis Riedel, 2019
Trigonopterus santubongensis Riedel, 2014
Trigonopterus sarinoi Narakusumo & Riedel, 2021
Trigonopterus sasak Riedel, 2014
Trigonopterus satu Riedel, 2014
Trigonopterus satyrus Riedel, 2019
Trigonopterus scabripes Riedel, 2019
Trigonopterus scabrosus Riedel, 2013
Trigonopterus scaphiformis Riedel, 2019
Trigonopterus scaphioides (Pascoe, 1871)
Trigonopterus scharfi Riedel, 2013
Trigonopterus schulzi Riedel, 2014
Trigonopterus scissops Riedel, 2013
Trigonopterus scitulus Riedel, 2019
Trigonopterus sculptirostris (Lea, 1928)
Trigonopterus sculpturatus Rheinheimer, 2013
Trigonopterus sebelas Riedel, 2014
Trigonopterus seclusus (Faust, 1899)
Trigonopterus sejunctus (Faust, 1899)
Trigonopterus selaruensis Narakusumo & Riedel, 2019
Trigonopterus selayarensis Riedel, 2019
Trigonopterus sellatus (Faust, 1899)
Trigonopterus sembilan Riedel, 2014
Trigonopterus semicribrosus Fairmaire, 1881
Trigonopterus semirubrus (Hustache, 1925)
Trigonopterus sepuluh Riedel, 2014
Trigonopterus seriatus Riedel, 2014
Trigonopterus serratifemur Riedel, 2014
Trigonopterus serratipes Marshall, 1931
Trigonopterus serripes Riedel, 2019
Trigonopterus seticnemis Riedel, 2019
Trigonopterus setifer Riedel, 2014
Trigonopterus setipoides Rheinheimer, 2013
Trigonopterus setipus Rheinheimer, 2013
Trigonopterus sexmaculatus Rheinheimer, 2013
Trigonopterus signicollis Riedel, 2013
Trigonopterus silaliensis Van Dam & Riedel, 2016
Trigonopterus silvestris Riedel, 2014
Trigonopterus silvicola Riedel, 2019
Trigonopterus similis (Faust, 1899)
Trigonopterus simulans Riedel, 2013
Trigonopterus singkawangensis Riedel, 2014
Trigonopterus singularis Riedel, 2014
Trigonopterus sinuatus Riedel, 2014
Trigonopterus soiorum Riedel, 2013
Trigonopterus solidus (Faust, 1899)
Trigonopterus sordidus Riedel, 2013
Trigonopterus squalidulus Riedel, 2019
Trigonopterus squalidus Riedel, 2014
Trigonopterus squamirostris Riedel, 2013
Trigonopterus squamosus (Lea, 1928)
Trigonopterus striatipennis (Lea, 1928)
Trigonopterus striatus Riedel, 2013
Trigonopterus strigatus Riedel, 2013
Trigonopterus strombosceroides Riedel, 2013
Trigonopterus subglabratus Riedel, 2013
Trigonopterus submetallicus Marshall, 1921
Trigonopterus subrubricollis Heller, 1920
Trigonopterus sulawesiensis Riedel, 2019
Trigonopterus sulcatus Riedel, 2013
Trigonopterus sumatrensis Riedel, 2014
Trigonopterus sumbawensis Riedel, 2014
Trigonopterus sundaicus Riedel, 2014
Trigonopterus sutrisnoi Narakusumo & Riedel, 2021
Trigonopterus suturaelevata Rheinheimer, 2013
Trigonopterus suturalis Riedel, 2014
Trigonopterus suturatus Riedel, 2019
Trigonopterus syarbis Riedel, 2014
Trigonopterus taenzleri Riedel, 2013
Trigonopterus talpa Riedel, 2013
Trigonopterus tanimbarensis Narakusumo & Riedel, 2019
Trigonopterus tanah Narakusumo & Riedel, 2021
Trigonopterus tatorensis Riedel, 2019
Trigonopterus taurekaorum Riedel, 2013
Trigonopterus tejokusumoi Narakusumo & Riedel, 2021
Trigonopterus telagensis Riedel, 2014
Trigonopterus tenuipes Riedel, 2019
Trigonopterus tepalensis Riedel, 2014
Trigonopterus terraereginae Riedel, 2016
Trigonopterus tialeorum Riedel, 2013
Trigonopterus tibialis Riedel, 2013
Trigonopterus tiga Riedel, 2014
Trigonopterus toboliensis Narakusumo & Riedel, 2021
Trigonopterus tolitoliensis Narakusumo & Riedel, 2021
Trigonopterus tomohonensis Riedel, 2019
Trigonopterus toraja Riedel, 2019
Trigonopterus tounaensis Narakusumo & Riedel, 2021
Trigonopterus tridentatus Riedel, 2013
Trigonopterus trigonopterus Riedel, 2014
Trigonopterus triradiatus Narakusumo & Riedel, 2019
Trigonopterus tujuh Riedel, 2014
Trigonopterus ujungkulonensis Riedel, 2014
Trigonopterus unicolor Rheinheimer, 2013
Trigonopterus uniformis Riedel, 2013
Trigonopterus unyil Narakusumo & Riedel, 2021
Trigonopterus vandekampi Riedel, 2010
Trigonopterus vanus (Faust, 1899)
Trigonopterus variabilis Riedel, 2013
Trigonopterus variolosus Riedel, 2014
Trigonopterus velaris Riedel, 2013
Trigonopterus verrucosus Riedel, 2013
Trigonopterus vicinus Riedel, 2019
Trigonopterus viduus Riedel, 2019
Trigonopterus violaceus Riedel, 2013
Trigonopterus viridescens Riedel, 2013
Trigonopterus volcanorum Riedel, 2019
Trigonopterus vulcanicus Riedel, 2014
Trigonopterus wallacei Riedel, 2014
Trigonopterus wamenaensis Riedel, 2013
Trigonopterus wangiwangiensis Riedel, 2019
Trigonopterus wariorum Riedel, 2013
Trigonopterus watsoni Riedel, 2019
Trigonopterus yoda Riedel, 2019
Trigonopterus yorkensis Riedel, 2016
Trigonopterus zonatus Heller, 1920
Trigonopterus zygops Riedel, 2013

Gallery

The lateral aspect on thorax of the weevil Trigonopterus oblongus. In this genus, the metanepisternite is absent and the elytron touches the metaventrite (indicated).

Metatarsus of weevil Trigonopterus sp. (from Java); scanning electron micrograph

Diagram of the biological screw in the weevil Trigonopterus oblongus

References

Riedal, Alexander (27 July 2011). "The weevil genus Trigonopterus Fauvel (Coleoptera, Curculionidae) and its synonyms—a taxonomic study on the species tied to its genus-group names". Zootaxa. 2977. doi:10.11646/zootaxa.2977.1.1.
Riedel, A.; Sagata, K.; Surbakti, S.; Tänzler, R.; Balke, M. (2013). "One hundred and one new species of Trigonopterus weevils from New Guinea". ZooKeys (280): 1–150. doi:10.3897/zookeys.280.3906. PMC 3677382. PMID 23794832.
Alexander Riedel (2010). "One of a thousand - a new species of Trigonopterus (Coleoptera, Curculionidae, Cryptorhynchinae) from New Guinea". Zootaxa. 2403: 59–68. doi:10.11646/zootaxa.2403.1.5.
Riedel, A.; Sagata, K.; Suhardjono, Y. R.; Tänzler, R.; Balke, M. (2013). "Integrative taxonomy on the fast track - towards more sustainability in biodiversity research". Frontiers in Zoology. 10 (1): 15. doi:10.1186/1742-9994-10-15. PMC 3626550. PMID 23537182.
Alexander Riedel, Daawia Daawia & Michael Balke (2010). "Deep cox1 divergence and hyperdiversity of Trigonopterus weevils in a New Guinea mountain range (Coleoptera, Curculionidae)". Zoologica Scripta. 39 (1): 63–74. doi:10.1111/j.1463-6409.2009.00404.x. S2CID 84957334.
Riedel, Alexander; Tänzler, Rene (21 January 2016). "Revision of the Australian species of the weevil genus Trigonopterus Fauvel". ZooKeys (556): 97–162. doi:10.3897/zookeys.556.6126. PMC 4740874. PMID 26877696.
Sheikh, Knvul (21 January 2016). "Hiding in Plain Sight: 24 New Beetle Species Discovered in Australia". Live Science. Retrieved 21 January 2016.
Van Dam, Matthew H.; Laufa, Raymond; Riedel, Alexander (21 April 2016). "Four new species of Trigonopterus Fauvel from the island of New Britain (Coleoptera, Curculionidae)". ZooKeys (582): 129–141. doi:10.3897/zookeys.582.7709. PMC 4857049. PMID 27199589.
Riedel, Alexander; Narakusumo, Raden Pramesa (7 March 2019). "One hundred and three new species of Trigonopterus weevils from Sulawesi". ZooKeys (828): 1–153. doi:10.3897/zookeys.828.32200. PMC 6418079. PMID 30940991.
Narakusumo, Raden Pramesa; Riedel, Alexander (22 October 2021). "Twenty-eight new species of Trigonopterus Fauvel (Coleoptera, Curculionidae) from Central Sulawesi". ZooKeys (1065): 29–79. doi:10.3897/zookeys.1065.71680.
van de Kamp, T.; Cecilia, A.; dos Santos Rolo, T.; Vagovič, P.; Baumbach, T.; Riedel, A. (2015). "Comparative thorax morphology of death-feigning flightless cryptorhynchine weevils (Coleoptera: Curculionidae) based on 3D reconstructions". Arthropod Structure & Development. 44 (6): 509–523. doi:10.1016/j.asd.2015.07.004. PMID 26259678.
Van De Kamp, T.; Dos Santos Rolo, T.; Vagovič, P.; Baumbach, T.; Riedel, A. (2014). "Three-Dimensional Reconstructions Come to Life – Interactive 3D PDF Animations in Functional Morphology". PLOS ONE. 9 (7): e102355. doi:10.1371/journal.pone.0102355. PMC 4100761. PMID 25029366.
Ross, Valerie (30 June 2011). "Zooming in on Beetles' Knees, Biologists Find Tiny Screws and Nuts". Discover magazine. Kalmbach Publishing Co. Retrieved 22 May 2011.
van de Kamp, Thomas; Vagovic, Patrik; Baumbach, Tilo; Riedel, Alexander (1 July 2011). "A Biological Screw in a Beetle's Leg". Science. 333 (6038): 52. doi:10.1126/science.1204245. PMID 21719669. S2CID 8527127.
Brown, Mark (5 July 2011). "Weevil Has Nuts and Bolts in Its Legs". Wired Science. Condé Nast Digital. Retrieved 25 July 2011.
Karlsruhe Institute of Technology (source) (5 July 2011). "Nature uses screws and nuts: Previously unknown musculoskeletal system discovered in weevils". ScienceDaily. Retrieved 25 July 2011.