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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Coleopterida
Ordo: Coleoptera
Subordo: †Protocoleoptera
Superfamilia: †Tshekardocoleoidea

Familia: †Tshekardocoleidae
Genera (13): AvocoleusBoscoleusEocoleusMoravocoleus – Permocoleus – ProsperocoleusRetelytronSylvacoleodesSylvacoleusTshekardocoleusUmoricoleusUralocoleusVotocoleus

Name

Tshekardocoleidae Rohdendorf, 1944
Synonyms

Uralocoleidae Zalessky, 1947
Moravocoleidae Kukalová-Peck & Beutel, 2012 [syn. Kirejtshuk et al., 2013]

References
Primary references

Kirejtshuk, A.G., Poschmann, M., Prokop, J., Garrouste, R. & Nel, N. 2014. [Online 2013] Evolution of the elytral venation and structural adaptations in the oldest Palaeozoic beetles (Insecta: Coleoptera: Tshekardocoleidae). Journal of Systematic Palaeontology 12(5): 575–600. DOI: 10.1080/14772019.2013.821530 Paywall. Reference page.
Kukalová-Peck, J. & Beutel, R.G. 2012. Is the Carboniferous †Adiphlebia lacoana really the "oldest beetle"? Critical reassessment and description of a new Permian beetle family. European Journal of Entomology 109(4): 633–645. DOI: 10.14411/eje.2012.075 Open access. Reference page.

Additional references

Bouchard, P., Bousquet, Y., Davies, A.E., Alonso-Zarazaga, M.A., Lawrence, J.F., Lyal, C.H.C., Newton, A.F., Reid, C.A.M., Schmitt, M., Ślipiński, S.A. & Smith, A.B.T. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88: 1–972. DOI: 10.3897/zookeys.88.807 Open access. PMC: 3088472 Open access. Reference page.
Kirejtshuk, A.G. & Nel, A. 2016. Происхождение отряда жуков (Coleoptera) и значение палеонтологической летописи [Origin of the Coleoptera and significance of the fossil record]. Euroasian Entomological Journal 15(Suppl. 1): 66–73. (in Russian). PDF. Reference page.

Links

The Paleobiology Database
Legalov, A.A. (coordinator): Tshekardocoleidae Species Listing in Joel Hallan's Biology Catalog
Tree of Life Web Project 2006. Tshekardocoleidae. Version 10 February 2006 (temporary) in The Tree of Life Web Project


Tshekardocoleidae is an exinct family of beetles, known from the Permian. Like other primitive beetles, they are thought to have been xylophagous.[1] The oldest known beetle, Coleopsis, was originally assigned to this family, but is now assigned to its own family Coleopseidae.[2] They represent some of the earliest known beetles, first appearing during the Cisuralian, before becoming extinct at the beginning of the Guadalupian.[1] A claimed Jurassic record is doubtful.[2]
Taxonomy
undescribed Tshekardocoleid larvae, Tshekarda site

Kirejtshuk (2020) included the following genera in Tshekardocoleidae[2]

Avocoleus Ponomarenko, 1969 - Obora site (Permian Sakmarian 295 to 290.1 Ma); Moravia, Czech Republic
Avocoleus fractus (Kukalová, 1969) (Type species)
Avocoleus neglegens (Kukalová, 1969)
Boscoleus Kukalová, 1969 - Obora site (Permian Sakmarian 295 to 290.1 Ma); Moravia, Czech Republic
Boscoleus blandus Kukalová, 1969 (Type species)
Eocoleus Kukalová, 1969 - Obora site (Permian Sakmarian 295 to 290.1 Ma); Moravia, Czech Republic
Eocoleus scaber Kukalová, 1969 (Type species)
Moravocoleus Kukalová, 1969 - Obora site (Permian Sakmarian 295 to 290.1 Ma); Moravia, Czech Republic
Moravocoleus permianus Kukalová, 1969 (Type species)
Permocoleus Lubkin & Engel, 2005 - Midco Member (Permian, Artinskian 290.1 to 279.3 Ma); Noble County, Oklahoma, USA
Permocoleus wellingtoniensis Lubkin & Engel, 2005 (Type species)
Prosperocoleus Kukalová, 1969 - Obora site (Permian Sakmarian 295 to 290.1 Ma); Moravia, Czech Republic
Prosperocoleus prosperus Kukalová, 1969 (Type species)
Retelytron Kukalová, 1965 - Obora site (Permian Sakmarian 295 to 290.1 Ma); Moravia, Czech Republic
Retelytron conopeum Kukalová, 1965 (Type species)
Sylvacoleodes Ponomarenko, 1963 - Tshekarda site (Permian Kungurian 295 to 290.1 Ma); Suksun, Perm Krai, Russia
Sylvacoleodes admirandus Ponomarenko, 1963 (Type species)
Sylvacoleus Ponomarenko, 1963 - Tshekarda site (Permian Kungurian 295 to 290.1 Ma); Suksun, Perm Krai, Russia
Sylvacoleus richteri Ponomarenko, 1963 (Type species)
Sylvacoleus sharovi Ponomarenko, 1963
Tshekardocoleus Rohdendorf, 1944 - Tshekarda site (Permian Kungurian 295 to 290.1 Ma); Suksun, Perm Krai, Russia
Tshekardocoleus magnus Rohdendorf, 1944 (Type species)
Tshekardocoleus minor Ponomarenko, 1963
Umoricoleus Kukalová, 1969 - Obora site (Permian Sakmarian 295 to 290.1 Ma); Moravia, Czech Republic
Umoricoleus perplex Kukalová, 1969 (Type species)
Votocoleus Kukalová, 1969 - Obora site (Permian Sakmarian 295 to 290.1 Ma); Moravia, Czech Republic
Votocoleus submissus Kukalová, 1969 (Type species)

The genus Uralocoleus had been placed in the family by some authors, but the type species Uralocoleus splendens Zalessky 1947 was considered incertae sedis by Kirejtshuk (2020) as a possible beetle or cockroach. The second species placed in the genus "Uralocoleus" ultimus was rejected from placement in Uralocoleus and listed as "genus incertus" ultimus Ponomarenko, 2000.[2]
References

Zhao, Xianye; Yu, Yilun; Clapham, Matthew E; Yan, Evgeny; Chen, Jun; Jarzembowski, Edmund A; Zhao, Xiangdong; Wang, Bo (2021-11-08). Perry, George H; Fikacek, Martin (eds.). "Early evolution of beetles regulated by the end-Permian deforestation". eLife. 10: e72692. doi:10.7554/eLife.72692. ISSN 2050-084X.
Kirejtshuk, Alexander G. (2020-02-17). "Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea". Geosciences. 10 (2): 73. Bibcode:2020Geosc..10...73K. doi:10.3390/geosciences10020073. ISSN 2076-3263.

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