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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Hexapoda
Classis: Insecta
Cladus: Dicondylia
Subclassis: Pterygota
Cladus: Metapterygota
Infraclassis: Neoptera
Cladus: Eumetabola
Cladus: Endopterygota
Superordo: Panorpida
Cladus: Amphiesmenoptera
Ordo: Lepidoptera
Subordo: Glossata
Cladus: Coelolepida
Cladus: Myoglossata
Cladus: Neolepidoptera
Infraordo: Heteroneura
Cladus: Eulepidoptera
Cladus: Ditrysia
Cladus: Apoditrysia
Cladus: Obtectomera
Superfamilia: Papilionoidea

Familia: Hesperiidae
Subfamilia: Eudaminae
Tribus: Eudamini
Subtribus: Loboclina
Genus: Zestusa
Species (4): Z. dorus – Z. elwesi – Z. levona – Z. staudingeri

Zestusa Lindsey, 1925

Type species: Plestia staudingeri Mabille, 1888.

Plestia Mabille, 1888: 146 (preocc. Plestia Stål, [1870] [Hemiptera]).
Type species: Plestia staudingeri Mabille, 1888.
Zestusa Lindsey, 1925: 106 (repl. Plestia Mabille, 1888).

Primary references

Lindsey, 1925, Ann. ent. Soc. Amer. 18.
Mabille, P. 1888. Description de Lépidoptéres (Hespérides) nouveaux. Le Naturaliste (10)2(31): 146–148, 169-171, 180-181. Reference page.

Additional references

Grishin, N.V. 2019. Expanded phenotypic diagnoses for 24 recently named new taxa of Hesperiidae (Lepidoptera). The Taxonomic Report 8(1): 1–15. PDF. Reference page.
Lamas, G. 2004. (ed.) Checklist: Part 4A. Hesperioidea - Papilionoidea. In Heppner, J.B. (ed.) Atlas of Neotropical Lepidoptera. Vol. 5A. Association for Tropical Lepidoptera/Scientific Publishers, Gainesville. 439p. Reference page.
Li, W., Cong. Q., Shen, J.H., Zhang, J., Hallwachs, W., Janzen, D.H. & Grishin, N.V. 2019. Genomes of skipper butterflies reveal extensive convergence of wing patterns. PNAS 116(13): 6232–6237. DOI: 10.1073/pnas.1821304116. Text. SI Appendix. Reference page.
Steinhauser, S.R. 1972. The genus Zestusa (Hesperiidae) in El Salvador with description of a new species. Journal of the Lepidopterists' Society 26(3): 127–132. PDF. Reference page.

Zestusa is a genus of skippers in the family Hesperiidae subfamily Eudaminae.

The genus occurs in Mexico and the western United States.

Natural History Museum Lepidoptera genus database

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