Mesolimulus

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Ecdysozoa
Cladus: Panarthropoda
Phylum: Arthropoda
Subphylum: Chelicerata
Classis: Merostomata
Ordo: Xiphosura
Subordo: Xiphosurida
Infraordo: Limulina

Familia: Limulidae
Genus: †Mesolimulus

Mesolimulus is an extinct genus of horseshoe crab. The best known examples are found in Solnhofen limestone near Solnhofen, Bavaria, Germany. Originally assigned to the living genus Limulus, they are related to and look virtually identical to modern horseshoe crabs. Other species assigned to Mesolimulus have been recorded spanning over 140 million years from the Middle Triassic to Late Cretaceous from England, Spain, Siberia and Morocco.
Kouphichnium
Main article: Kouphichnium
Mesolimulus walchi fossil and track

The unusual trackways left by these animals at Solnhofen were originally thought to have been made by birds or pterosaurs, since they included a cross-shaped marking comparable with the impression of a typical bird foot. Eventually they were correctly identified as arthropod footprints and named Kouphichnium walchi, as proved by some remarkable 'death traces' showing a Mesolimulus circling round on itself before dying. The last leg of more advanced horseshoe crabs is modified into a so-called 'pusher' which consists of four plates at the tip which push against the soft sediment rather like a snow-shoe. This was what left the unusual bird-like footprints.
Mesolimulus walchi

Horseshoe crabs in general date to the Ordovician Period, more than 440 million years ago, and late Paleozoic Euproops fossils indicate that they have changed little over the last 300 million years. Fossils preserved in Solenhofen limestone are unusual because soft body parts and skeletons are clearly represented.
Species

After[1]

Mesolimulus sibiricus Ponomarenko, 1985, Middle Jurassic (Bathonian) Talynzhansk Formation, Russia
Mesolimulus tafraoutensis Lamsdell et al., 2020 Late Cretaceous (upper Cenomanian –lower Turonian) Gara Sbaa Lagerstatte, Morocco
Mesolimulus walchi (Desmarest, 1822) Late Jurassic (Kimmeridgian –Early Tithonian) Solnhofen Limestone, Germany
Mesolimulus woodwardi (Watson, 1909) Middle Jurassic (Aalenian) Northampton Sand Formation, England
Mesolimulus crespelli Vıa Boada, 1987 Middle Triassic (Ladinian) Alcover Limestone Formation, Spain

References

Bicknell, Russell D. C.; Błażejowski, Błażej; Wings, Oliver; Hitij, Tomaž; Botton, Mark L. (2021-03-02). Zhang, Xi‐Guang (ed.). "Critical re‐evaluation of Limulidae uncovers limited Limulus diversity". Papers in Palaeontology. 7 (3): 1525–1556. doi:10.1002/spp2.1352. ISSN 2056-2799. S2CID 233783546.

Caster, K. E. 1941. Trails of Limulus and supposed vertebrates from Solnhofen lithographic limestone. Pan-American Geologist, 76:241–258.
Malz, H. 1964. Kouphichnium walchi, die Geschichte einer Fährte und ihres

Tieres. Natur und Museum, 94:81–97.