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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Amphibia
Subclassis: Lissamphibia
Ordo: Anura

Familia: Microhylidae
Subfamilia: Asterophryinae
Genus: Cophixalus
Species (55): C. aenigma – C. aimbensis – C. albolineatus – C. amabilis – C. ateles – C. australis – C. balbus – C. bewaniensis – C. biroi – C. bombiens – C. cateae – C. caverniphilus – C. cheesmanae – C. clapporum – C. concinnus – C. crepitans – C. cryptotympanum – C. cupricarenus – C. daymani – C. desticans – C. exiguus – C. hannahae – C. hinchinbrookensis – C. hosmeri – C. humicola – C. infacetus – C. interruptus – C. iovaorum – C. kaindiensis – C. kethuk – C. kulakula – C. linnaeus – C. mcdonaldi – C. melanops – C. misimae – C. monosyllabus – C. montanus – C. monticola – C. neglectus – C. nexipus – C. nubicola – C. ornatus – C. pakayakulangun – C. parkeri – C. peninsularis – C. petrophilus – C. phaeobalius – C. pictus – C. pipilans – C. pulchellus – C. rajampatensis – C. riparius – C. salawatiensis – C. saxatilis – C. shellyi – C. sisyphus – C. sphagnicola – C. tagulensis – C. tenuidactylus – C. tetzlaffi – C. timidus – C. tomaiodactylus – C. tridactylus – C. variabilis – C. verecundus – C. verrucosus – C. viridis – C. wempi – C. zweifeli
Name

Cophixalus Boettger, 1892

Type species: Sphenophryne verrucosa Boulenger, 1898
Synonyms

Cophixalus Boettger, 1892
Phrynixalus Boettger, 1895

References

Boettger 1892, Kat. Batr. Samml. Mus. Senckenb. Naturforsch. Ges.: 24.
Günther, R. 2003: First record of the microhylid frog genus Cophixalus from western Papua, Indonesia, with descriptions of two new species (Anura: Microhylidae). Herpetozoa, 16(1-2): 3–21. [not seen]
Günther, R. 2010: Another new Cophixalus species (Amphibia: Anura: Microhylidae) from western New Guinea. Bonn zoological Bulletin, 57(2): 231–240. PDF
Günther, R., Richards, S.J., Tjaturadi, B., Krey, K. 2015. Two new species of the genus Cophixalus from the Raja Ampat Islands west of New Guinea (Amphibia, Anura, Microhylidae). Zoosystematics and Evolution 91(2): 199-213. DOI: 10.3897/zse.91.5411 Open access. Reference page.
Hoskin, C.J. 2004: Australian microhylid frogs (Cophixalus and Austrochaperina): phylogeny, taxonomy, calls, distributions and breeding biology. Australian journal of zoology, 52(3): 237–269. DOI: 10.1071/ZO03056 PDF
Hoskin, C.J. 2013: A new frog species (Microhylidae: Cophixalus) from boulder-pile habitat of Cape Melville, north-east Australia. Zootaxa 3722(1): 61–72. DOI: 10.11646/zootaxa.3722.1.5 Reference page.
Hoskin, C.J.; Aland, K. 2011: Two new frog species (Microhylidae: Cophixalus) from boulder habitats on Cape York Peninsula, north-east Australia. Zootaxa, 3027: 39–51. Preview
Kraus, F. 2012. Papuan frogs of the genus Cophixalus (Anura: Microhylidae): new synonyms, new species, and a dichotomous key. Zootaxa 3559: 1–36. PreviewPaywall Reference page.
Kraus, F.; Allison, A. 2009: New species of Cophixalus (Anura: Microhylidae) from Papua New Guinea. Zootaxa, 2128: 1–38. Abstract & excerpt
Kraus, F.; Allison, A. 2009: New microhylid frogs from the Muller Range, Papua New Guinea. ZooKeys, 26: 53–76. DOI: 10.3897/zookeys.26.258

Links

Amphibian Species of the World: an Online Reference. Version 6.0. Cophixalus access date 14 July 2015
ION [needs update]
Nomenclator Zoologicus

Vernacular names

Cophixalus (rainforest frogs or nursery frogs) is a genus of microhylid frogs.[citation needed] These are arboreal species with expanded toe-pads, endemic to Moluccan Islands, New Guinea and northeastern Queensland, Australia.[citation needed]
Species

As of December 2019, Amphibian Species of the World assigns 67 species to the genus Cophixalus.[1]

The source column gives direct links to the sources used:

IUCN description of species at International Union for Conservation of Nature Red List of Threatened Species. IUCN Red List categories are:

EX - Extinct, EW - Extinct in the Wild
CR - Critically Endangered, EN - Endangered, VU - Vulnerable
NT - Near Threatened, LC - Least Concern
DD - Data Deficient, NE - Not Evaluated

ASW description of species at Amphibian Species of the World.

Binomial name and authority Common name Geographic range CS Source
Cophixalus aenigma Hoskin, 2004 Thornton rainforest frog north-eastern Queensland VU ASW IUCN
Cophixalus albolineatus Kraus, 2012 Mount Shungol, New Guinea ASW
Cophixalus amabilis Kraus, 2012 Woodlark Island (PNG) ASW
Cophixalus ateles (Boulenger, 1898) Papua rainforest frog Mount Obree (New Guinea, PNG); possibly widespread in south-eastern Papua New Guinea DD ASW IUCN
Cophixalus australis Hoskin, 2012[2] southern ornate nursery-frog north-east Queensland ASW
Cophixalus balbus Günther, 2003 Yapen rainforest frog Yapen Island (Indonesia) DD ASW IUCN
Cophixalus bewaniensis Kraus & Allison, 2000 Menawa rainforest frog Bewani Mountains (New Guinea, PNG) DD ASW IUCN
Cophixalus biroi (Méhely, 1901) Sattelberg rainforest frog northern New Guinea (Indonesia and Papua New Guinea) LC ASW IUCN
Cophixalus bombiens Zweifel, 1985 buzzing rainforest frog north-eastern Queensland NT ASW IUCN
Cophixalus cateae Richards and Günther, 2019 Agogo Range in Southern Highlands Province, Papua New Guinea ASW
Cophixalus caverniphilus Kraus and Allison, 2009 Muller Range (New Guinea, PNG) ASW
Cophixalus cheesmanae Parker, 1934 Kokoda rainforest frog eastern Papua New Guinea LC ASW IUCN
Cophixalus clapporum Kraus, 2012 Woodlark Island (PNG) ASW
Cophixalus concinnus Tyler, 1979 rattling rainforest frog north-eastern Queensland (Australia) CR ASW IUCN
Cophixalus crepitans Zweifel, 1985 rusty rainforest frog eastern Cape York Peninsula (Queensland, Australia) NT ASW IUCN
Cophixalus cryptotympanum Zweifel, 1956 Zweifel's rainforest frog Mount Dayman (New Guinea, PNG) DD ASW IUCN
Cophixalus cupricarenus Kraus and Allison, 2009 Rossel Island (PNG) ASW
Cophixalus daymani Zweifel, 1956 Dayman rainforest frog Mount Dayman (PNG) DD ASW IUCN
Cophixalus desticans Kraus and Allison, 2009 south-easternmost tip of New Guinea, Normanby Island (PNG) ASW
Cophixalus exiguus Zweifel & Parker, 1969 Cooktown rainforest frog north-eastern Queensland (Australia) NT ASW IUCN
Cophixalus hannahae Richards and Günther, 2019 central cordillera of Papau New Guinea ASW
Cophixalus hinchinbrookensis Hoskin, 2012[2] Hinchinbrook Island nursery-frog Hinchinbrook Island (Queensland, Australia) ASW
Cophixalus hosmeri Zweifel, 1985 clicking rainforest frog northern Queensland (Australia) LC ASW IUCN
Cophixalus humicola Günther, 2006 Waira rainforest frog Yapen Island, northern New Guinea (Indonesia) LC ASW IUCN
Cophixalus infacetus Zweifel, 1985 chocolate rainforest frog north-eastern Queensland (Australia) LC ASW IUCN
Cophixalus interruptus Kraus and Allison, 2009 Morobe Province (New Guinea, PNG) ASW
Cophixalus iovaorum Kraus and Allison, 2009 Mount Obree (New Guinea, PNG) ASW
Cophixalus kaindiensis Zweifel, 1979 Kaindi rainforest frog Mount Kaindi and Mount Missim (New Guinea, PNG) DD ASW IUCN
Cophixalus kethuk Kraus and Allison, 2009 Rossel Island (PNG) ASW
Cophixalus kulakula Hoskin and Aland, 2011 north-eastern Queensland (Australia) ASW
Cophixalus linnaeus Kraus and Allison, 2009 Bowutu Mountains (New Guinea, PNG) ASW
Cophixalus mcdonaldi Zweifel, 1985 Mcdonald's rainforest frog north-eastern Queensland (Australia) EN ASW IUCN
Cophixalus melanops Kraus and Allison, 2009 Tagula Island (PNG) ASW
Cophixalus misimae Richards & Oliver, 2007 Misima rainforest frog Misima Island (PNG) CR ASW IUCN
Cophixalus monosyllabus Günther, 2010 southern Fakfak Mountains (New Guinea, Indonesia) ASW
Cophixalus montanus (Boettger, 1895) mountain rainforest frog Halmahera Island (Indonesia) DD ASW IUCN
Cophixalus monticola Richards, Dennis, Trenerry, & Werren, 1994 carbine rainforest frog northern Queensland (Australia) EN ASW IUCN
Cophixalus neglectus Zweifel, 1962 brown rainforest frog northern Queensland (Australia) EN ASW IUCN
Cophixalus nexipus Kraus, 2012 Mount Obree (New Guinea, PNG) ASW
Cophixalus nubicola Zweifel, 1962 Michael rainforest frog Mount Michael (New Guinea, PNG) VU ASW IUCN
Cophixalus ornatus (Fry, 1912) ornate rainforest frog mid-eastern Queensland, Hinchinbrook (Australia) LC ASW IUCN
Cophixalus pakayakulangun Hoskin and Aland, 2011 north-eastern Queensland (Australia) ASW
Cophixalus parkeri Loveridge, 1948 Parker's rainforest frog central mountainous region of Papua New Guinea LC ASW IUCN
Cophixalus peninsularis Zweifel, 1985 peninsula rainforest frog northern Queensland (Australia) DD ASW IUCN
Cophixalus petrophilus Hoskin, 2013 blotched boulder-frog [3]
Cophixalus phaeobalius Kraus and Allison, 2009 Bowutu Mountains (New Guinea, PNG) ASW
Cophixalus pictus Kraus, 2012 West Papua Province (New Guinea, Indonesia) ASW
Cophixalus pipilans Zweifel, 1980 Sempi rainforest frog from Lae to the Adelbert Mountains (New Guinea, PNG) LC ASW IUCN
Cophixalus pulchellus Kraus & Allison, 2000 pretty rainforest frog Mount Hunstein (New Guinea, PNG) DD ASW IUCN
Cophixalus rajampatensis Günther, Richards, Tjaturadi, and Krey, 2015 Raja Ampat Island group off of western New Guinea (Indonesia) ASW
Cophixalus riparius Zweifel, 1962 Wilhelm rainforest frog central mountain ranges of Papua Guinea (PNG) LC ASW IUCN
Cophixalus salawatiensis Günther, Richards, Tjaturadi, and Krey, 2015 Raja Ampat Island group off of western New Guinea (Indonesia) ASW
Cophixalus saxatilis Zweifel & Parker, 1977 Black Mountain rainforest frog northern Queensland (Australia) VU ASW IUCN
Cophixalus shellyi Zweifel, 1956 Shelly's rainforest frog central mountain ranges of Papua New Guinea, Adelbert Mountains, Huon Peninsula LC ASW IUCN
Cophixalus sphagnicola Zweifel & Allison, 1982 Morobe rainforest frog Mount Amungwiwa, Mount Kaini, and Mount Missim (New Guinea, PNG) LC ASW IUCN
Cophixalus tagulensis Zweifel, 1963 Tagula rainforest frog Tagula Island (PNG) DD ASW IUCN
Cophixalus tenuidactylus Günther and Richards, 2012 Kaijende Highlands (New Guinea, PNG) ASW
Cophixalus tetzlaffi Günther, 2003 Bomberai rainforest frog Bomberai Peninsula (New Guinea, Indonesia) DD ASW IUCN
Cophixalus timidus Kraus & Allison, 2006 Simpson rainforest frog Mount Simpson (New Guinea, PNG) DD ASW IUCN
Cophixalus tomaiodactylus Kraus and Allison, 2009 Bowutu Mountains (New Guinea, PNG) ASW
Cophixalus tridactylus Günther, 2006 Wondiwoi rainforest frog Wondiwoi Mountains (New Guinea, Indonesia) DD ASW IUCN
Cophixalus variabilis Kraus & Allison, 2006 Pekopekowana rainforest frog south-eastern Papua New Guinea LC ASW IUCN
Cophixalus verecundus Zweifel & Parker, 1989 Myola rainforest frog Mount Bellamy (New Guinea, PNG); presumably more widely distributed DD ASW IUCN
Cophixalus verrucosus (Boulenger, 1898) Moroke rainforest frog southern and eastern Papua New Guinea LC ASW IUCN
Cophixalus viridis Günther, Richards, and Dahl, 2014 Green rainforest frog Muller Range, Western Province, Papua New Guinea ASW
Cophixalus wempi Richards and Oliver, 2010 Southern Highlands Province (New Guinea, PNG) ASW
Cophixalus zweifeli Davies & McDonald, 1998 Cape Melville rainforest frog northern Queensland (Australia) DD ASW IUCN

References

Frost, Darrel R. 2019. Amphibian Species of the World: an Online Reference. Version 6.0 (10 December 2019). Cophixalus[permanent dead link]. Electronic Database accessible at http://research.amnh.org/herpetology/amphibia/index.php. American Museum of Natural History, New York, USA. (Accessed: November 17, 2013).
Hoskin, Conrad J. (2012). "Two new frog species (Microhylidae: Cophixalus) from the Australian Wet Tropics region, & redescription of Cophixalus ornatus" (PDF). Zootaxa. 3271: 1–16. doi:10.11646/zootaxa.3271.1.1.
Conrad J. Hoskin (2013). "A new frog species (Microhylidae: Cophixalus) from boulder-pile habitat of Cape Melville, north-east Australia" (PDF excerpt). Zootaxa. 3722 (1): 61–72. doi:10.11646/zootaxa.3722.1.5. PMID 26171515.

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