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Superregnum: Eukaryota
Supergroup: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Classis: Amphibia
Subclassis: Lissamphibia
Ordo: Anura

Familia: Centrolenidae
Subfamilia: Centroleninae
Tribus: Cochranellini
Genus: Rulyrana
Species (8): R. adiazeta – R. erminea – R. flavopunctata – R. mcdiarmidi – R. saxiscandens – R. spiculata – R. susatamai – R. tangarana
Name

Rulyrana Guayasamin, Castoviejo-Fisher, Trueb, Ayarzagüena, Rada & Vilà, 2009
References

Guayasamin, J.M; Castroviejo-Fisher, S.; Trueb, L.; Ayarzagüena, J.; Rada, M.; Vilà, C. 2009: Phylogenetic systematics of glassfrogs (Amphibia: Centrolenidae) and their sister taxon Allophryne rithveni. Zootaxa, 2100: 1–97. Abstract & excerpt PDF

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Rulyrana is a small genus of glass frogs.[2][3] They are found in South America, on the Amazonian slopes of the Andes in Ecuador, Peru, and possibly Bolivia, as well as on the eastern slopes of the Cordillera Central and the western slopes of the Cordillera Oriental in Colombia.[2]

Etymology

The generic name Rulyrana honors Pedro Ruiz-Carranza [fr] and John D. Lynch [fr] who have "contributed enormously to the understanding of centrolenid diversity, biology, and evolution". The name is made up from the two first letters of their surnames in combination with rana for frog. In addition, "Ruly" is the nickname of Martín Bustamante, who has also contributed to amphibian conservation.[1]
Description

Rulyrana have moderate to extensive webbing between the third and fourth fingers. The dorsum is lavender in preserved individuals and may have spots. Internal features include green bones (in live specimens), lobed liver that is covered by a transparent hepatic peritoneum, whereas the ventral parietal peritoneum is white in its anterior part and transparent in its poster part. The digestive tract is translucent. In terms of osteology, Rulyrana have dentigerous process in the vomer and—usually—vomerine teeth. The humeral spines are not present.[1]

While distinct from most other glass frogs, there are no characters that could unambiguously place a species in Rulyrana or in the genus Sachatamia; genetic data are needed for an unambiguous allocation. The two genera, however, have largely disjunct distribution areas (Rulyrana are found in the Amazon Basin and the Colombian Cordillera Central while Sachatamia are not found further east than the Colombian Cordillera Central).[1]
Reproduction

The males call while sitting on upper sides of leaves or rocks. The eggs are deposited on leaves or rocks.[1]
Species

There are six species:[2]

Rulyrana adiazeta (Ruiz-Carranza and Lynch, 1991)
Rulyrana flavopunctata (Lynch and Duellman, 1973)
Rulyrana mcdiarmidi (Cisneros-Heredia, Venegas, Rada, and Schulte, 2008)
Rulyrana saxiscandens (Duellman and Schulte, 1993)
Rulyrana spiculata (Duellman, 1976)
Rulyrana susatamai (Ruiz-Carranza and Lynch, 1995)

The AmphibiaWeb includes also Sachatamia orejuela in this genus.[3]
References

Guayasamin, J. M.; Castroviejo-Fisher, S.; Trueb, L.; Ayarzagüena, J.; Rada, M.; Vilà, C. (2009). "Phylogenetic systematics of glassfrogs (Amphibia: Centrolenidae) and their sister taxon Allophryne ruthveni". Zootaxa. 2100: 1–97. doi:10.11646/zootaxa.2100.1.1.
Frost, Darrel R. (2017). "Rulyrana Guayasamin, Castroviejo-Fisher, Trueb, Ayarzagüena, Rada, and Vilà, 2009". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 6 May 2017.
"Centrolenidae". AmphibiaWeb. University of California, Berkeley. 2017. Retrieved 6 May 2017.

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