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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Subsectio: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Theropoda
Cladus: Neotheropoda
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Euornithes
Cladus: Ornithuromorpha
Cladus: Ornithurae
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Ordo: Charadriiformes
Subordo: Alcae

Familia: Alcidae
Genus: Cepphus
Species: C. carbo – C. columba – C. grylle

Cepphus Pallas, 1769

Spicilegia zoologica 1 fasc.5 p. 33

Vernacular names
suomi: Riskilät

Cepphus is a genus of seabirds in the auk family also referred to as true guillemots or, in North America, simply as guillemots. The genus name Cepphus is from Ancient Greek κέπφος kepphos,[1] a pale waterbird mentioned by Greek authors including Aristotle.[2] The English word "guillemot" is from French guillemot probably derived from Guillaume, "William".[3] "Murre" is of uncertain origins, but may imitate the call of the common guillemot.[4]

These are medium-sized birds with mainly black plumage in the breeding season, thin dark bills and red legs and feet. Two species have white wing patches, the third has white facial “spectacles”. They are much paler in winter plumage, mottled above and white below.

The breeding habitat is rocky shores and islands on the coasts of the northern Atlantic and Pacific Oceans. They usually lay their eggs in rocky sites near water.

These birds may overwinter in their breeding areas, moving to open waters if necessary, but usually not migrating very far south.

They dive for food from the surface, swimming underwater. They mainly eat fish and crustaceans, also some molluscs, insects and plant material.

The species are:

Image Scientific name Common Name Distribution
Gryllteiste 5.JPG Cepphus grylle Black guillemot or Tystie Circumpolar: Arctic coasts, North Atlantic, Alaska
Pigeon Guillemot at Living Coasts.jpg Cepphus columba Pigeon guillemot Kuril Islands and the Kamchatka Peninsula in Siberia to coasts in western North America from Alaska to California.
SpectacledGuillemot.jpg Cepphus carbo Spectacled guillemot Sea of Okhotsk and the Kuril Islands in Russia and on the northern island of Hokkaidō in Japan

There are also fossil forms

Cepphus olsoni Howard, 1982 (San Luis Rey River Late Miocene - Early Pliocene of W USA)
Cepphus storeri Harrison, 1977 (Red Crag of Suffolk Late Miocene - England
Cepphus cf. columba (Lawrence Canyon Early Pliocene of W USA)
Cepphus cf. grylle (San Diego Late Pliocene, W USA)

The latter two resemble the extant species, but because of the considerable distance in time or space from their current occurrence may represent distinct species.

κέπφος. Liddell, Henry George; Scott, Robert; A Greek–English Lexicon at the Perseus Project.
Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 96. ISBN 978-1-4081-2501-4.
"Guillemot". Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription or participating institution membership required.)
"Murre". Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription or participating institution membership required.)

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