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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Subsectio: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Cladus: Theropoda
Cladus: Neotheropoda
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Euornithes
Cladus: Ornithuromorpha
Cladus: Ornithurae
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Ordo: Strigiformes

Familia: Strigidae
Subfamilia: Striginae
Genus: Ketupa
Species: K. flavipes - K. ketupu - K. zeylonensis
Name

Ketupa Lesson, 1830: 114
References

Lesson, R.P. 1830–1831. Traité d'ornithologie, ou, Tableau méthodique des ordres, sous-ordres, familles, tribus, genres, sous-genres et races d'oiseaux : ouvrage entièrement neuf, formant le catalogue le plus complet des espèces réunies dans les collections publiques de la France. F. G. Levrault, Paris. Vol. 1: pp. i–xxxii, 1–659. BHL Reference page. [original description: p. 114]

Fish owls form a group of owls in the family Strigidae.[1] Ketupa was proposed as generic name by René-Primevère Lesson in 1831 for fish owl species from Java and India.[2] Results of a phylogenetic analysis of nine horned owl species indicate that Ketupa species form a monophyletic group.[3] Ketupa is therefore thought to merit subgeneric rank comprising the following species:[1]

Brown fish owl (K. zeylonensis) described by Johann Friedrich Gmelin in 1788 was a fish owl from Sri Lanka;[4]
Buffy fish owl (K. ketupu) described by Thomas Horsfield in 1821 was a fish owl from Java;[5]
Tawny fish owl (K. flavipes) described by Brian Houghton Hodgson in 1836 was a fish owl from Nepal.[6]

Blakiston's fish owl (Bubo blakistoni) described by Henry Seebohm in 1883 is a fish owl from Japan and the Vladivostok region of Russia. It was thought to form the link between Ketupa and Bubo species, as its toes are bare of feathers like in Ketupa species, but it is much larger.[7] Since its first description, and placement in the genus Bubo, it has repeatedly been placed in Ketupa; the 2013 phylogenetic analysis of nine horned owl species indicate that four Ketupa species including Blakiston's fish owl form a monophyletic group,[3] and genetic testing of Blakiston's fish owl in Russia published in 2018 indicates a fairly large divergence from the genomes of typical Bubo.[8]


References

del Hoyo, J.; Collar, N. J.; Christie, D. A.; Elliott, A.; Fishpool, L. D. C. (2014). "Strigidae". In del Hoyo, J.; Elliott, A.; Sargatal, J.; Christie, D. A.; de Juana, E. (eds.). Handbook of the Birds of the World and BirdLife International Illustrated Checklist of the Birds of the World. Vol. 1: Non-passerines. Barcelona, Spain and Cambridge, UK: Lynx Edicions and BirdLife International.
Lesson, R.-P. (1831). "Sous-genre. Ketupu; Ketupa". Traité d'ornithologie, ou, Tableau méthodique des ordres, sous-ordres, familles, tribus, genres, sous-genres et races d'oiseaux : ouvrage entièrement neuf, formant le catalogue le plus complet des espèces réunies dans les collections publiques de la France. Vol. 1. Paris: F. G. Levrault. p. 114.
Omote, K.; Nishida, C.; Dick, M. H.; Masuda, R. (2013). "Limited phylogenetic distribution of a long tandem-repeat cluster in the mitochondrial control region in Bubo (Aves, Strigidae) and cluster variation in Blakiston's fish owl (Bubo blakistoni)". Molecular Phylogenetics and Evolution. 66 (3): 889–897. doi:10.1016/j.ympev.2012.11.015. PMID 23211719.
Gmelin, J. F. (1788). "Strix zeylonensis". Caroli a Linné, Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Vol. I (13th aucta, reformata ed.). Lipsiae: Georg Emanuel Beer. p. 287.
Horsfield, T. (1821). "Systematic Arrangement and Description of Birds from the Island of Java". Transactions of the Linnean Society. 13 (1): 133–200. doi:10.1111/j.1095-8339.1821.tb00061.x.
Hodgson, B. H. (1836). "On a new Piscatory Genus of the Strigine Family". Journal of the Asiatic Society of Bengal. 5: 363–365.
Seebohm, H. (1883). "Exhibition of, and remarks upon, a new Owl (Bubo blakistoni) from Yezo". Proceedings of the Zoological Society of London (20 November): 466.
Spiridonova, L. N.; Surmach, S. G. (2018). "Whole mitochondrial genome of Blakiston's Fish Owl Bubo (Ketupa) blakistoni suggests its redescription in the genus Ketupa". Russian Journal of Genetics. 54 (3): 369–373. doi:10.1134/S1022795418030134.

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