Milvus milvus

Milvus milvus (*)

Cladus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Aves
Subclassis: Carinatae
Infraclassis: Neornithes
Parvclassis: Neognathae
Ordo: Falconiformes
Familia: Accipitridae
Subfamilia: Milvinae
Genus: Milvus
Species: Milvus milvus
Subspecies: M. m. fasciicauda - M. m. milvus

Name

Milvus milvus (Linnaeus, 1758)

Reference

Systema Naturae ed.10 p.89

Vernacular names
Internationalization
Català: Milà reial
Česky: Luňák červený
Ελληνικά : Ψαλιδιάρης
English: Red Kite
Español: Milano real
Nederlands: Rode wouw
Türkçe: Kızıl çaylak
Українська: Шуліка рудий

The Red Kite (Milvus milvus) is a medium-large bird of prey in the family Accipitridae, which also includes many other diurnal raptors such as eagles, buzzards, and harriers. The species is currently endemic to the Western Palearctic region in Europe and northwest Africa, though formerly also occurred just outside in northern Iran.[2] It is a rare species which is resident in the milder parts of its range in western Europe and northwest Africa, but birds from northeastern and central Europe winter further south and west, reaching south to Turkey. Vagrants have reached north to Finland and south to Israel and Libya.[2]

Taxonomy

This species was first described by Linnaeus in his Systema naturae in 1758 as Falco milvus.[3]

The Red Kite has been known to successfully hybridize with the Black Kite in captivity where both species were kept together, and in the wild on the Cape Verde Islands.[4] The Red Kites on the Cape Verde Islands are (or rather were) quite distinct in morphology, being somewhat intermediate with Black Kites. The question whether the Cape Verde Kite should be considered a distinct species (Milvus fasciicauda) or a Red Kite subspecies has not been settled. A recent mtDNA study on museum specimens suggests that Cape Verde birds did not form a monophyletic lineage among or next to Red Kites.[5]

However, this interpretation is problematic: mtDNA analysis is very susceptible to hybridization events, the evolutionary history of the Cape Verde population is not known, and the genetic relationship of Red Kites in general is very confusing, with geographical proximity being no indicator of genetic relatedness and the overall genetic similarity high,[6] perhaps indicating a relict species.

Given the morphological distinctness of the Cape Verde birds and the fact that the Cape Verde population was isolated from other populations of Red Kites, it cannot be conclusively resolved at this time whether the Cape Verde population was not a distinct subspecies (as M. migrans fasciicauda) or even species that frequently absorbed stragglers from the migrating European populations into its gene pool. More research seems warranted, but at any rate the Cape Verde population is effectively extinct since 2000, all surviving birds being hybrids with Black Kites (which merely raises further questions about their taxonomic status).[5]

Description

The Red Kite is 60–66 cm (24-27 in) long with a 175–195 cm wingspan; males have a weight of 800–1200 g, and females 1000–1300 g.[2] It is an elegant bird, soaring with long wings held at a dihedral, and long forked tail twisting as it changes direction. The body, upper tail and wing coverts are rufous. The white primary flight feathers contrast with the black wing tips and dark secondaries. Apart from the weight difference, the sexes are similar, but juveniles have a buff breast and belly. The call is a thin piping, similar to but less mewling than the Common Buzzard. There is also a rare white leucistic form which accounts for approximately 1% of hatchlings in the Welsh population but is at a disadvantage in the survival stakes.[7]

Differences between adults and juveniles

Adults differ from juveniles in a number of characteristics:

Adults are overall more deeply rufous, compared with the more washed out colour of juveniles;
Adults have black breast-streaks whereas on juveniles these are pale;
Juveniles have a less deeply-forked tail, with a dark subterminal band;
Juveniles have pale tips to all of the greater-coverts (secondary and primary) on both the upper- and under-wings, forming a long narrow pale line; adults have pale fringes to upperwing secondary-coverts only.

These differences hold throughout most of the first year of a bird's life.

Behaviour


The Red Kite's diet consists mainly of small mammals such as mice, voles, shrews, young hares and rabbits. It also feeds on a wide variety of carrion including sheep carcasses and dead game birds. Live birds are also taken and occasionally reptiles and amphibians. Earthworms also form an important part of the diet, especially in the spring.[8]

As scavengers, red kites are particularly sensitive to poisoning.[9] Illegal poison baits set for foxes or crows are indiscriminate and kill protected birds and other animals.

At signs of danger a mother will signal the young who will "play dead" to the extent that a fox will believe them to be dead and leave them, thinking it can return to eat them later.[10]

Breeding

Adult red kites are sedentary birds, and they occupy their breeding home range all year in the United Kingdom, though many of the global populations are migratory (particularly the Swedish population, which winters in Spain).[11] Each nesting territory can contain up to five alternative nest sites. Both birds build the nest on a main fork or a limb high in a tree, 12-20m high made of dead twigs and lined with grass or other vegetation and sheep’s wool.[8][11]

Distribution and status


The Red Kite inhabits broadleaf woodlands, valleys and wetland edges, to 800 m. It is endemic to the western Palearctic, with the European population of 19,000-25,000 pairs encompassing 95% of its global breeding range. It breeds from Spain and Portugal east through central Europe to Ukraine, north to southern Sweden, Latvia and the UK, and south to southern Italy. There is also a population in northern Morocco. Northern birds move south in winter, mostly staying in the west of the breeding range, but also to eastern Turkey, northern Tunisia and Algeria. The three largest populations (in Germany, France and Spain, which together hold more than 75% of the global population) all declined during 1990-2000, and overall the species declined by almost 20% over the ten years. The main threats to this species are poisoning, through illegal direct poisoning and indirect poisoning due to pesticides, particularly in the wintering ranges in France and Spain, and changes in agricultural practices causing a reduction in food resources. Other threats include electrocution, hunting and trapping, deforestation, egg-collection (on a local scale) and possibly competition with the generally more successful Black Kite M. migrans.[1]
[edit] Continental Europe

German populations declined by 25-30% between 1991 and 1997, but have remained stable since then, with the populations of the northern foothills of the Harz Mountains (the most densely populated part of its range) suffering an estimated 50% decline from 1991-2001. In Spain the species showed an overall decline in breeding population of up to 43% for the period 1994 to 2001-02, and surveys of wintering birds in 2003-04 suggest a similarly large decline in core wintering areas. The Balearic Islands population has declined from 41-47 breeding pairs in 1993 to just 10 in 2003. In France, breeding populations have decreased in the northeast, but seem to be stable in southwest and central France and Corsica. However, populations elsewhere are stable or undergoing increases. In Sweden the species has increased from 30-50 pairs in the 1970s to 1,200 breeding pairs in 2003. In Switzerland, populations increased during the 1990s, and have now stabilised.[1] However, according to a report by the Welsh Kite Trust, the UK is the only country in which the Red Kite population is increasing. Red Kites are decreasing in their three strongholds of Spain, France and Germany.[12]

United Kingdom

In the United Kingdom Red Kites were once so common that William Shakespeare described London as " a city of Red Kites and Crows". As ubiquitous scavengers they lived on carrion and garbage, giving rise to their common name in Elizabethan England - Shitehawk. Shakespeare's King Lear describes his daughter Goneril as a detested kite, and he also wrote "when the kite builds, look to your lesser linen" in reference to them stealing washing that was hung out to dry during the nesting season. [13] In the mid 15th century King James II of Scotland decreed that they should be "killed wherever possible", but they remained protected in England and Wales for the next 100 years as they kept the streets free of carrion and rotting food.[14] Under Tudor "vermin laws" many creatures were seen as competitors for the produce of the countryside and bounties were paid by the parish for their carcasses.[15]

By the 20th century the breeding population had become restricted to a handful of pairs in Wales, but recently the Welsh population has been supplemented by re-introductions in England and Scotland. In 2004, from 375 occupied territories identified at least 216 pairs were thought to have hatched eggs and 200 pairs reared at least 286 young.[1] In 1989 six Swedish birds were released at a site in north Scotland and four Swedish and one Welsh bird in Buckinghamshire.[16] Altogether, 93 birds of Swedish and Spanish origin were released at each of the sites. In the second stage of reintroduction in 1995 and 1996, further birds were brought over from Germany to populate the areas of Dumfries and Galloway, and the Derwent Valley in North East England.[16]In Northern Ireland four pairs were released and in 2010 the first reproduction was recorded. The reintroductions in The Chilterns have been a particular success. Between 1989 and 1993 90 birds were released in the Chilterns Area of Outstanding Natural Beauty and by 2002, 139 pairs were breeding in the area.[17] Another very successful reintroduction has been in Northamptonshire, which has become a stronghold for the Red Kite[18]. Thirty Spanish birds were reintroduced into Rockingham Forest near Corby in 2000 [19], and by 2010, the RSPB estimated that over 200 chicks had been reared from that initial release. So successful has the reintroduction been that 30 chicks have been transported from Rockingham Forest, for release in Cumbria. [20]

A sighting of the first Red Kite to be seen in London for 150 years was reported in The Independent newspaper in January 2006[21] and in June of that year, the UK-based Northern Kites Project reported that kites had bred in the Derwent Valley in and around Rowlands Gill, Tyne and Wear for the first time since the re-introduction.[22]

In 1999 the Red Kite was named 'Bird of the Century' by the British Trust for Ornithology.[13]

In June 2010 the Forestry Commission North West England announced a three-year project that will see 90 Red Kites released in Grizedale Forest, Cumbria under a special licence issued by Natural England. The Grizedale programme will be the ninth reintroduction of Red Kites into different regions of the UK, and the final re-introduction phase in England.[23]

The stated aims of the Grizedale project are:

To establish a viable population of Red Kites in Grizedale, South Cumbria by 2015.
To increase the rate of Red Kite expansion into North West England and link up with existing populations in Wales, Yorkshire, North East England and South West Scotland and so increase the chances of a continuous geographical range.*To develop community involvement and create educational opportunities arising from the project.[24]

Ireland

Red Kites were extinct in Ireland by the middle nineteenth century, due to persecution, poisoning and woodland clearance. In May 2007, Minister for the Environment, Heritage and Local Government Dick Roche announced an agreement to bring at least 100 birds from Wales to restock the population as part of a 5-year programme in the Wicklow Mountains, similar to the earlier Golden Eagle reintroduction programme.[25] On the 19 July 2007 the first thirty red kites were released in Co. Wicklow.[26][27] On 22 May 2010, 2 newly-hatched Red Kite chicks were discovered in the Wicklow mountains,[28] bringing the number of chicks hatched since reintroduction to 7.[29]
Populations and trends by country

The following figures (mostly estimates) have been collated from various sources.[2][30][31][32][33][34] They cover most of the countries in which Red Kites are believed to have bred.
Country Year Pairs Trend Notes
Norway 1980 0 0 Bred occasionally in 19th century
Sweden late 2000s c.1800 + Increase from 400 pairs in 1993
Denmark 2009 75–80[35] + Extinct c.1920, then recolonised (from Sweden) 1970s
Scotland 2009 135 + Extinct 1886, reintroduced 1989–1992
Northern Ireland 2007 4 + First successful breeding reported in 2010 following reintroduction in 2008
Ireland 2010 7 + First successful breeding reported in 2010 following reintroduction in 2007
Wales 2009 c.1000 + Declined to 2 pairs in 1930s, then recovery
England 2011 c.2000 + Extinct 1870s, reintroduced 1989–1992
France late 2000s c.3000 – 2300–2900 pairs 1980s
Netherlands c.1998 <5 + Extinct 1852, recolonised 1976
Belgium c.1995 50–60 + Declined to 1–3 pairs early 1970s, then recovery
Luxembourg 1997 46 +
Germany late 2000s c.12000 – 15000–25000 pairs 1980s
Poland c.1998 650–700 + 400–450 pairs 1980s
Estonia 1989 <1 ?
Latvia 1992 0–50 + Extinct 1964, then recolonised
Lithuania 1988 1–2 + Extinct, then recolonised 1981
Russia 1992 0–50 ?
Belarus 1997 1 +/– Extinct 1950s, recolonised 1985; 10 pairs 1990
Ukraine 1990 5–8 –
Czech Republic 1993–94 30–50 + Extinct late 19th century, recolonised 1975
Slovakia 1992 10–20 ?–
Switzerland c.1995 800–1000 + Declined 19th century, later recovery; 235–300 pairs late 1980s
Austria 2000 0–2 – Extinct 1950, recolonised 1970s; 10 pairs 1990
Hungary c.1998 1+ – 30 pairs 1950s
Romania 1995 15–20 ?+
Moldova 1990 1 ?
Bulgaria 0 ? May breed but no proof
Croatia 0 ? 2–5 pairs 1980s
Bosnia and Herzegovina 0 ?
Yugoslavia 0 ? Formerly commoner
Montenegro 1995 0 ?
Serbia ?
Macedonia 0 ?
Italy c.2002 300–400 0/+ 70–150 pairs late 1980s
Albania ? Bred 1906
Greece 0 ?
Turkey 0 ? May have bred in past but no proof
Algeria 0 ? Bred in 19th century, now extinct
Tunisia 0 ? Bred in 19th century, now extinct
Spain late 2000s c.2200 – 10000 pairs 1977
Portugal c.1995 100–200 +/0
Morocco c.1992 10–100 – In danger of extinction
Canary Islands 0 0 Extinct 1970s
Cape Verde 2000 1? – 50–75 pairs late 1980s; effectively extinct
Observation

One of the best places to see the Red Kite in Scandinavia is Scania in southern Sweden. It may be observed in one of its breeding locations such as the Kullaberg Nature Preserve near Molle.[36]

Some of the best places to see them in the UK are Gigrin Farm near Rhayader, mid Wales, where hundreds are fed by the local farmer as a tourist attraction.[37] and the nearby Nant-Yr-Arian forest recreation centre in Ceredigion[38] where the rare leucistic variant can be seen.[39] In the UK the Oxfordshire part of the Chilterns has many Red Kites.[17] They can also be seen around Harewood near Leeds where they were re-introduced in 1999.[40]

References

^ a b c d Anon (2009). "Red Kite - BirdLife Species Factsheet". 2009 IUCN Red List Category. BirdLife International. Retrieved 2009-07-05.
^ a b c d Snow, D. W. & Perrins, C. M. (1998). The Birds of the Western Palearctic Concise Edition. OUP ISBN 0-19-854099-X.
^ (Latin) Linnaeus, C (1758). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.. Holmiae. (Laurentii Salvii).. pp. 89. "F. cera flava, cauda forsicata, corpore ferrugineo, capite albidiore.."
^ Sabine Hille and Jean-marc Thiollay (2000). The imminent extinction of the Kites Milvus milvus fasciicauda and Milvus m. migrans on the Cape Verde Islands. Bird Conservation International, 10 , pp 361-369
^ a b Johnson,1,2*, Jeff; Watson R, Mindell D. (7 july 2005). "Prioritizing species conservation: does the Cape Verde kite exist?". Proc Biol Sci. (The Royal Society) 272 (7): 1365–1371. doi:10.1098/rspb.2005.3098. PMC 1560339. PMID 16006325. Retrieved 26 October 2009.
^ Schreiber, Arnd; Stubbe, Michael & Stubbe, Annegret (2000): Red kite (Milvus milvus) and black kite (M. migrans): minute genetic interspecies distance of two raptors breeding in a mixed community (Falconiformes: Accipitridae). Biol. J. Linn. Soc. 69(3): 351–365. doi:10.1006/bijl.1999.0365 (HTML abstract)
^ Anon. "The White Kite". Gigrin Farm - The Red Kite feeding station. Gigrin Farm. Retrieved 2009-07-07.[dead link]
^ a b Pugh, Effyn. "The Red Kite". bitdsofbritain.co.uk. Birds of Britain. Retrieved 2009-10-26.
^ "Wildlife crime soars". The Herald Series. 12 September 2007. Retrieved 6 July 2009. "In Didcot, poisoned rabbits were laid out as bait disguised as road-kill, targeting red kites"
^ Anon. "Red Kite". Animal Corner. Animal Corner. Retrieved 26 october 2009.
^ a b Anon. "Scotland's Wildlife: Red Kite". BBC Scotland outdoors articles. BBC. Retrieved 26 October 2009.
^ "Red Kites decline in Europe". Welsh Kite Trust. undated. Retrieved 2007-02-24.
^ a b BBC Radio 4, Debating Animals, Series 2, The Kestrel and Red Kite by Rod Liddle
^ Atrill, Rod. "The Red Kite in West Wales". New Key on Cardigan Bay in West Wales. Rod Attrill. Retrieved 2009-07-07.
^ McCarthy, Michael (23 March 2007). "Book Review:Silent Field, By Roger Lovegrove:songbirds versus shotguns". The Independent: (Independent.co.uk). Retrieved 2009-07-07.
^ a b Anon. "Red Kite". RSPB Conservation. RSPB. Retrieved 2009-10-26.
^ a b Schurmer, Michael (November 2002). "Breeding Bird Survey of the Chilterns Area of Outstanding Natural Beauty 2002". RSPB. Retrieved 26 October 2009.
^ "RSPB Red Kite numbers are soaring across the UK". Birdguides. Monday 15th September 2008. Retrieved 4 August 2011.
^ "Red Kite project a soaring success". Evening Telegraph. Friday 9 November 2007. Retrieved 4 August 2011.
^ "Red kite chicks from Northamptonshire released to wild". BBC. 17 August 2010. Retrieved 4 August 2011.
^ McCarthy, Michael (13 january 2006). "Shakespeare's red kite returns to London after an absence of 150 years". The Independent on Sunday (Independent News and Media Limited). Retrieved 2009-07-05.
^ "Delight as red kite chicks hatch". BBC News. 16 June 2006. Retrieved 2006-10-27.
^ Anon (17 June 2010). "Red kites to make a Lake District return". The Westmorland Gazette (Newsquest media group). Retrieved 12 July 2010.
^ Anon. "Grizedale Red Kite Project". Forstry Commission information posters. Forestry commission. Retrieved 12 July 2010.
^ Anon. "Golden Eagle Trust, Glenveagh National Park". National Development plan. NDP. Retrieved 26 october 2009.
^ News - Department of the Environment, Heritage & Local Government
^ Anon. "Red Kites fly again in Ireland.". Wildlife Extra: News. Wildlife Extra. Retrieved 26 October 2009.
^ http://www.independent.ie/national-news/two-chicks-about-the-size-of-a-fist-2190451.html
^ http://www.goldeneagle.ie/news_viewnews.php?x=3&z=44&news_id=9&article=277
^ Carter, Ian (2001): The Red Kite. Arlequin Press, Chelmsford, UK. 187pp.
^ Cramp, S. (1980). The Birds of the Western Palearctic Volume 2. Oxford ISBN 0-19-857505-X.
^ Holloway, S. (1996). The Historical Atlas of Breeding Birds in Britain and Ireland: 1875–1900. T & A D Poyser ISBN 0-85661-094-1.
^ RSPB Scotland, cited in The Scotsman, Monday 15th September 2008
^ Carter, Ian (2009): The Red Kite. presentation to the Cambridgeshire Bird Club, Cambridge, UK, 13 November 2009.
^ Gert Hjembæk (2009-11-25). "Den røde drage bliver hængende i Danmark". Dansk Ornitologisk Forening. Retrieved 2009-11-28.
^ Hogan, C. Michael (2005): Kullaberg Nature Reserve, Sweden. Lumina technologies.
^ "Red Kite Feeding Station — Gigrin Farm". Retrieved 2006-10-27.
^ Anon (2008). "Bwlch Nant-yr-Arian". Bwlch Nant Yr Arian Visitor Centre. Retrieved 2009-07-07.
^ Melton, Tom (13 August 2008). "Leucistic Red Kite". ephotozone. ephotozone. Retrieved 2009-07-07.
^ Anon. "Birder watchers' paradise". BBC Hands on Nature:. BBC. pp. Parks: Harewood Estate. Retrieved 26 october 2009.

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