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Cladus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Classis: Aves
Subclassis: Carinatae
Infraclassis: Neornithes
Parvclassis: Neognathae
Ordo: Passeriformes
Subordo: Passeri
Parvordo: Passerida
Superfamilia: Sylvioidea
Familia: Sittidae
Genera: Sitta - Tichodroma


Sittidae (Lesson, 1828)

Vernacular names
Česky: Brhlíkovití
한국어: 동고비과

Sittidae is a family of small passerine birds which contains the single genus Sitta containing about 24 species of nuthatches, which are found across Eurasia and North America.

Sometimes the Wallcreeper (Tichodroma muraria), which is restricted to the mountains of southern Eurasia, is also placed in this family, in a separate subfamily "Tichodromadinae", in which case the nuthatches are classified in the subfamily "Sittinae". However, it is more often placed in a separate family, the Tichodromadidae.[1] The Wallcreeper is intermediate in its morphology between the nuthatches and the treecreepers, but its appearance, the texture of its plumage, and the shape and pattern of its tail suggest that it is closer to the former taxon.[2]


The Sittidae family was described by René-Primevère Lesson in 1828. Its closest relatives, other than the Wallcreeper, are the treecreepers, and the (two or) three families are sometimes placed in a larger grouping with the wrens and gnatcatchers. This superfamily, the Certhioidea is proposed on phylogenetic studies using mitochondrial and nuclear DNA, and was created to cover a clade of (four or) five families removed from a larger grouping of passerine birds, the Sylvioidea.[3]

The Nuthatch Vanga of Madagascar and the sitellas from Australia and New Guinea were once placed in the family Sittidae, because of similarities in appearance and lifestyle to the nuthatches, but they are not closely related, the resemblances arising from convergent evolution to fill an ecological niche.[4]



All the species in this group nest in cavities. The Western and Eastern Rock Nuthatches use rock crevices, and the rest of the nuthatches nest in tree holes. The chicks are altricial, which means they are blind, featherless and helpless at birth. Both parents feed the nestlings until the young birds fledge.


Invertebrates are a major part of the diet for nuthatches, especially during the breeding season, but most species also eat seeds at least during the winter, when invertebrates are less readily available. Larger food items, such as big insects, snails, acorns or seeds may be wedged into cracks and hacked with the nuthatches's strong bill, this of course being the behaviour which gives that subfamily group its name.[4] The nuthatches all store food, usually seeds, which may be pushed into crevices or into the ground, hidden under small stones, or tucked behind bark flakes; the rock nuthatches will also wedge snails into suitable crevices for consumption in times of need. Caches are recovered by memory,[5] and can be retrieved as long as 98 days after being stored.[6] In one study of European Nuthatches birds refrained from using their caches during benign conditions in order to save them for harsher conditions.


The fossil record for this group appears to be restricted to a foot bone of an early Miocene bird from Bavaria which has been identified as an extinct representative of the climbing Certhioidea, a clade comprising the treecreepers, Wallcreeper and nuthatches. It has been described as Certhiops rummeli.[7]


Most nuthatches have large populations and extensive geographical ranges, and present few conservation problems.[8]

A few of the more restricted nuthatch species are threatened by deforestation.

* The White-browed Nuthatch is endangered. The population of a few thousand birds is decreasing, and no conservation measures are in place.[9][10]

* The endangered Algerian Nuthatch occurs only at four known sites in Algeria, and it is possible that the total population does not exceed 1,000 birds.[11]

* The Yunnan Nuthatch is vulnerable, although still locally common.[12]

* The Yellow-billed Nuthatch is vulnerable especially on Hainan, where more than 70% of the woodland has been lost in 50 years.[13]

* The Krüper's Nuthatch is near-threatened in its stronghold in Turkey, where urbanisation and development for tourism are placing considerable pressure on mature coniferous forest.[14]


1. ^ Snow, David; Perrins, Christopher M (editors) (1998). The Birds of the Western Palearctic (BWP) concise edition (2 volumes). Oxford: Oxford University Press. pp. 1408–1410. ISBN 019854099X.
2. ^ Vaurie, Charles; Koelz, Walter, (November 1950). "Notes on some Asiatic nuthatches and creepers" (PDF). American Museum novitates 1472: 1–39. http://digitallibrary.amnh.org/dspace/bitstream/2246/4241/1/N1472.pdf.
3. ^ Cracraft, J.; Barker, F. Keith; Braun, M. J.; Harshman, J.; Dyke, G.; Feinstein, J.; Stanley, S.; Cibois, A.; Schikler, P.; Beresford, P.; García-Moreno, J.; Sorenson, M. D.; Yuri, T.; Mindell. D. P. (2004) "Phylogenetic relationships among modern birds (Neornithes): Toward an avian tree of life." p468–489 in Assembling the tree of life (J. Cracraft and M. J. Donoghue, eds.). Oxford University Press, New York. ISBN 0195172345
4. ^ a b Harrap, Simon; Quinn, David (1996). Tits, Nuthatches and Treecreepers. Christopher Helm. pp. 16–17. ISBN 0-7136-3964-4.
5. ^ Hardling, Roger; Kallander, Hans & Jan-Åke Nilsson (1997). "Memory for Hoarded Food: An Aviary Study of the European Nuthatch" (PDF). The Condor 99 (2): 526–529. doi:10.2307/1369961. JSTOR 1369961. http://elibrary.unm.edu/sora/Condor/files/issues/v099n02/p0526-p0529.pdf.
6. ^ Nilsson, Jan-Åke; Persson, Hans Källander Owe (1993). "A prudent hoarder: effects of long-term hoarding in the European nuthatch, Sitta europaea". Behavioral Ecology 4 (4): 369–373. doi:10.1093/beheco/4.4.369.
7. ^ Manegold, Albrecht (April 2008). "Earliest fossil record of the Certhioidea (treecreepers and allies) from the early Miocene of Germany". Journal of Ornithology 149 (2): 223–228. doi:10.1007/s10336-007-0263-9.
8. ^ "Sitta". Species Search Results. BirdLife International. 2. Retrieved 2008-06-21.
9. ^ BirdLife International (2004). Sitta victoriae. 2007 IUCN Red List of Threatened Species. IUCN 2007. Retrieved on 18 June 2008.
10. ^ "Ecology of the White-browed Nuthatch Sitta victoriae in Natmataung National Park, Myanmar, with notes on other significant species" (PDF). Forktail 19: 57–62. 2003. http://www.orientalbirdclub.org/publications/forktail/19pdfs/Naing-Nuhatch.pdf.
11. ^ BirdLife International (2004). Sitta ledanti. 2007 IUCN Red List of Threatened Species. IUCN 2007. Retrieved on 17 June 2008.
12. ^ BirdLife International (2004). Sitta yunnanensis. 2007 IUCN Red List of Threatened Species. IUCN 2007. Retrieved on 18 June 2008.
13. ^ BirdLife International (2004). Sitta solangiae. 2007 IUCN Red List of Threatened Species. IUCN 2007. Retrieved on 18 June 2008.
14. ^ BirdLife International (2005). Sitta krueperi. 2007 IUCN Red List of Threatened Species. IUCN 2007. Retrieved on 17 June 2008.

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Source: WIkipedia, Wikispecies: All text is available under the terms of the GNU Free Documentation License