Stercorariidae

Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Archosauromorpha
Cladus: Crurotarsi
Divisio: Archosauria
Cladus: Avemetatarsalia
Cladus: Ornithodira
Subtaxon: Dinosauromorpha
Cladus: Dinosauriformes
Cladus: Dracohors
Cladus: Dinosauria
Ordo: Saurischia
Cladus: Eusaurischia
Subordo: Theropoda
Cladus: Neotheropoda
Cladus: Averostra
Cladus: Tetanurae
Cladus: Avetheropoda
Cladus: Coelurosauria
Cladus: Tyrannoraptora
Cladus: Maniraptoromorpha
Cladus: Maniraptoriformes
Cladus: Maniraptora
Cladus: Pennaraptora
Cladus: Paraves
Cladus: Eumaniraptora
Cladus: Avialae
Infraclassis: Aves
Cladus: Euavialae
Cladus: Avebrevicauda
Cladus: Pygostylia
Cladus: Ornithothoraces
Cladus: Ornithuromorpha
Cladus: Carinatae
Parvclassis: Neornithes
Cohors: Neognathae
Cladus: Neoaves
Ordo: Charadriiformes
Subordo: Lari

Familia: Stercorariidae
Genus: Stercorarius
Name

Stercorariidae Gray, 1871
Vernacular names
беларуская: Паморнікавыя
čeština: Chaluhovití
Deutsch: Raubmöwen
hrvatski: Pomornici
Türkçe: Korsan martıgiller ya da Yırtıcı martıgiller

The skuas /ˈskjuːə/ are a group of predatory seabirds with seven species forming the genus Stercorarius, the only genus in the family Stercorariidae. The three smaller skuas, the long-tailed jaeger, the parasitic jaeger, and the pomarine jaeger are called jaegers in American English. also spelled Yeager in some translations.[citation needed]

The English word "skua" comes from the Faroese name for the great skua, skúgvur [ˈskɪkvʊɹ], with the island of Skúvoy renowned for its colony of that bird. The general Faroese term for skuas is kjógvi [ˈtʃɛkvɪ]. The word "jaeger" is derived from the German word Jäger, meaning "hunter".[1][2] The genus name Stercorarius is Latin and means "of dung";[note 1] because the food disgorged by other birds when pursued by skuas was once thought to be excrement.[3]

Skuas nest on the ground in temperate and Arctic regions, and are long-distance migrants. They have even been sighted at the South Pole.[4]

Biology and habits
Two brown skuas (S. antarcticus) and a southern giant petrel (Macronectes giganteus) fighting over a dead Antarctic fur seal

Outside the breeding season, skuas take fish, offal, and carrion. Many practice kleptoparasitism, which comprises up to 95% of the feeding methods of wintering skuas, by chasing gulls, terns and other seabirds to steal their catches, regardless of the size of the species attacked (up to three times heavier than the attacking skua). The larger species, such as the great skua, also regularly kill and eat adult birds, such as puffins and gulls, and have been seen killing birds as large as a grey heron.[5] On the breeding grounds, the three, more slender northern breeding species commonly eat lemmings. Those species that breed in the southern oceans largely feed on fish that can be caught near their colonies. The eggs and chicks of other seabirds, primarily penguins, are an important food source for most skua species during the nesting season.[6]

In the southern oceans and Antarctica region, some skua species (especially the south polar skua) will readily scavenge carcasses at breeding colonies of both penguins and pinnipeds. Skuas will also kill live penguin chicks. In these areas, the skuas will often forfeit their catches to the considerably larger and very aggressive giant petrels. Skuas have also been observed to directly pilfer milk from the elephant seal's teat.[7]

They are medium to large birds, typically with grey or brown plumage, often with white markings on the wings.[6] The skuas range in size from the long-tailed skua, Stercorarius longicauda, at 310 grams (0.68 pounds), to the brown skua, Stercorarius antarcticus, at 1.63 kg (3.6 lb). On average, a skua is about 56 cm (22 in) long, and 121 cm (48 in) across the wings. They have longish bills with a hooked tip, and webbed feet with sharp claws. They look like large dark gulls, but have a fleshy cere above the upper mandible.

The skuas are strong, acrobatic fliers. They are generally aggressive in disposition. Potential predators approaching their nests will be quickly attacked by the parent birds, which usually target the heads of intruders – a practice known as 'divebombing'.[8]
Great skua leaving the nest
Skua nestling, with egg tooth still present on its beak
Taxonomy

The genus Stercorarius was introduced by the French zoologist Mathurin Jacques Brisson in 1760 with the parasitic jaeger (Stercorarius parasiticus) as the type species.[9][10]

Skuas are related to gulls, waders, auks, and skimmers. In the three smaller species, all nesting exclusively in the Holarctic, breeding adults have the two central tail feathers obviously elongated, and at least some adults have white on the underparts and pale yellow on the neck. These characteristics are not shared by the larger species, all native to the Southern Hemisphere except for the great skua. Therefore, the skuas are often split into two genera, with only the smaller species retained in Stercorarius, and the large species placed in Catharacta. However, based on genetics, behavior, and feather lice, the overall relationship among the species is best expressed by placing all in a single genus.[11] The pomarine and great skuas' mitochondrial DNA (inherited from the mother) is in fact more closely related to each other than it is to either Arctic or long-tailed skuas, or to the Southern Hemisphere species.[12] Thus, hybridization must have played a considerable role in the evolution of the diversity of Northern Hemisphere skuas.
Species

The genus contains seven species:[13]

Image	Scientific name	Common Name	Distribution
	Stercorarius chilensis	Chilean skua	Breeds along the coasts of southern Chile and southern Argentina, winters along the Pacific coasts of Peru and Chile as well as the Atlantic coast of Argentina
	Stercorarius maccormicki	South polar skua	Breeds along the coast on Antarctica, winters in the north Atlantic and north Pacific
	Stercorarius antarcticus	Brown skua	Southern Ocean
	Stercorarius skua	Great skua	Breeds along the coastline of the northeast Atlantic, winters in the north Atlantic
	Stercorarius pomarinus	Pomarine jaeger or pomarine skua	Breeds along the Arctic coastline, winters in tropical and subtropical oceans
	Stercorarius parasiticus	Parasitic jaeger or Arctic skua	Breeds along the Arctic coastline, winters in the southern hemisphere
	Stercorarius longicaudus	Long-tailed jaeger or long-tailed skua	Breeds in the Arctic, winters in the Southern Ocean

References

The word stercorārius is from stercus ("dung"), which is also the etymon of stercoranism, stercobilin, stercoral, etc.

"Jaeger". Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription or participating institution membership required.)
"Skua". Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription or participating institution membership required.)
Jobling, James A (2010). The Helm Dictionary of Scientific Bird Names. London: Christopher Helm. p. 365. ISBN 978-1-4081-2501-4.
Mark Sabbatini, "Non-human life form seen at Pole", The Antarctic Sun, 5 January 2003.
Scottish Ornithologists' Club Archived 2011-07-16 at the Wayback Machine
Harrison, Colin J.O. (1991). Forshaw, Joseph (ed.). Encyclopaedia of Animals: Birds. London: Merehurst Press. p. 109. ISBN 1-85391-186-0.
"Antarctica's Milk-Stealing, Grudge-Holding Seabirds". Boston NPR. 2016.
"Scottish Wildlife Trust builds £50,000 loo on Handa". BBC News. 12 March 2012.
Brisson, Mathurin Jacques (1760). Ornithologie, ou, Méthode Contenant la Division des Oiseaux en Ordres, Sections, Genres, Especes & leurs Variétés (in French and Latin). Paris: Jean-Baptiste Bauche. Vol. 1, p. 56, Vol. 6, p. 149.
Peters, James Lee, ed. (1934). Check-list of Birds of the World. Vol. 2. Cambridge, Massachusetts: Harvard University Press. p. 311.
American Ornithologists' Union (2000). Forty-second supplement to the American Ornithologists' Union Check-list of North American Birds. The Auk 117(3):847–858.
Cohen, Baker, Belchschmidt, Dittmann, Furness, Gerwin, Helbig, de Korte, Marshall, Palma, Peter, Ramli, Siebold, Willcox, Wilson and Zink (1997). Enigmatic phylogeny of skuas. Proc. Biol. Sci. 264(1379):181–190.
Gill, Frank; Donsker, David, eds. (2019). "Noddies, gulls, terns, auks". World Bird List Version 9.2. International Ornithologists' Union. Retrieved 24 June 2019.