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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
OrdoTherapsida
Cladus: Theriodontia
Subordo: Cynodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohort: Theria
Cohort: Eutheria
Cohort: Placentalia
Cladus: Boreoeutheria
Superordo: Euarchontoglires
Ordo: Primates
Subordo: Haplorhini
Infraordo: Simiiformes
Parvordo: Platyrrhini

Familia: Atelidae
Subfamilia: Alouattinae
Genus: Alouatta
Species: Alouatta coibensis
Name

Alouatta coibensis Thomas, 1902
Vernacular names
English: Coiba Island Howler
español: Mono Aullador de Coiba
português: Guariba-da-Ilha-Coiba
Türkçe: Coiba uluyan maymunu

The Coiba Island howler (Alouatta coibensis) is a type of howler monkey, a type of New World monkey, endemic to Panama. Although the Coiba Island howler has been recognized as a separate species by a number of authorities since a 1987 study of its fingerprints, mitochondrial DNA testing found it does not differ from mantled howler populations in any significant way.[4] A reason given for treating it as a separate species is that the dermal ridges of its hands and feet differ from those of the mantled howler.[5]

A. c. coibensis is smaller than other Central American howler monkeys and has duller pelage than the Azuero howler, Alouatta coibensis trabeata.
Subspecies

Two subspecies of this howler have been recognized by those who consider it a separate species:[1]

Alouatta coibensis coibensis Thomas, 1902, found on Coiba Island and Jicaron, off the Pacific coast of Panama
Azuero howler, A. c. trabeata Lawrence, 1933, endemic to the Azuero Peninsula in Panama.

References

Groves, C. P. (2005). Wilson, D. E.; Reeder, D. M. (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Baltimore: Johns Hopkins University Press. pp. 148–149. ISBN 0-801-88221-4. OCLC 62265494.
Cuarón, A. D.; Shedden, A.; Rodríguez-Luna, E.; de Grammont, P. C. & Link, A. (2008). "Alouatta palliata ssp. coibensis". IUCN Red List of Threatened Species. 2008. Retrieved 19 January 2012.
"Appendices | CITES". cites.org. Retrieved 2022-01-14.
Rylands; Groves; Mittermeier; Cortes-Ortiz & Hines (2006). "Taxonomy and Distributions of Mesoamerican Primates". In Estrada, A.; Garber, P.A.; Pavelka, M.S.M. & Luecke, L. (eds.). New Perspectives in the Study of Mesoamerican Primates. Springer. p. 53. ISBN 978-0-387-25854-6.
Rowe, N. (1996). The Pictorial Guide to the Living Primates. East Hampton, N.Y.: Pogonias Press. p. 109. ISBN 0-9648825-0-7.

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