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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Cladus: Cynodontia
Cladus: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Cladus: Boreoeutheria
Superordo: Laurasiatheria
Ordo: Chiroptera
Subordo: Yinpterochiroptera

Familia: Pteropodidae
Subfamilia: Cynopterinae
Tribus: Cynopterini
Genus: Cynopterus
Species: C. horsfieldii
Subspecies: C. h. harpax – C. h. horsfieldii – C. h. persimilis – C. h. princeps
Name

Cynopterus horsfieldii Gray, 1843

Type locality: Indonesia, Java.
References

Gray. 1843. List of Specimens Mammal Collection of the British Museum: 38.
Conservation status: IUCN: Cynopterus horsfieldii (Least Concern)
Cynopterus horsfieldii in Mammal Species of the World.
Wilson, Don E. & Reeder, DeeAnn M. (Editors) 2005. Mammal Species of the World – A Taxonomic and Geographic Reference. Third edition. ISBN 0-8018-8221-4.

Distribution

Thailand
Cambodia
West Malaysia
Borneo
Java
Sumatra
Lesser Sunda Isandls

Vernacular names
Deutsch: Horsfield-Kurznasenflughund
English: Horsfield's Fruit Bat

Horsfield's fruit bat (Cynopterus horsfieldii) is a species of megabat native to South East Asia. It is named for Thomas Horsfield, an American naturalist who presented the type specimen to the British Museum.
Description

Horsfield's fruit bat is a medium-sized megabat, intermediate in size between flying foxes and pygmy fruit bats. Adults weigh around 55 to 60 grams (1.9 to 2.1 oz), and have light-grey to brown fur, with a reddish-brown or orange mantle around the shoulders. In some males, the mantle extends across the chest, and the fur is often brighter in colour than in females. The rim of the ears and the skin overlying the metacarpals and phalanges within the wing are white. Juveniles have a more bland coat pattern, with uniformly dull buff or grey fur.[2]

The bats have a short, broad snout, ending in a pair of almost tubular nostrils. Both the eyes and ears are large, although the latter have a simpler structure than in most other bats, and lack tragi.[2] The wings have a low aspect ratio and high wing loading, typical of many megabats, and indicating a relatively slow flight speed and moderate manoeuvrability.[3]
Distribution and habitat

Horsfield's fruit bat is found in Thailand, Indonesia, Malaysia, and Brunei. Within this region, it inhabits a broad range of lowland habitats, from dense primary rain forest to agricultural land and suburban gardens.[1]

The four recognised subspecies are:

C. h. harpax - Thailand, peninsular Malaysia, Sumatra
C. h. horsfieldii - Java and islands east to Sumbawa
C. h. persimilis - Borneo
C. h. princeps - Nias

Biology and behaviour

Horsfield's fruit bats eat the fruit of strangler figs, Elaeocarpus, and Payena, and the flowers of bitter beans. They have been reported to pluck fruit from trees and carry it to roosts elsewhere to feed. During the dry season, when fruit is in short supply, they instead feed on pollen, which they take from a wide variety of plants.[2]

They live in small groups, consisting of a single adult male and up to five females and their young. Although these groups are maintained year-round, individual females often move between different groups, and may spend some time nesting alone between leaving one group and joining another. They roost in trees and cave mouths, reportedly favouring banana trees. They often modify their roosting sites by constructing tents from the leaves, partly cutting through them to make an inverted "V" shape.[4]

They breed throughout the year, but most commonly give birth at two times of the year, between February and March and between July and August. They have been reported to live for at least 31 months.[2]
References

Bates, P.; Francis, C.; Gumal, M.; Bumrungsri, S. (2019). "Cynopterus horsfieldii". IUCN Red List of Threatened Species. 2019: e.T6104A22113239. doi:10.2305/IUCN.UK.2019-3.RLTS.T6104A22113239.en. Retrieved 19 November 2021.
Campbell, P.; Kunz, T.H. (2006). "Cynopterus horsfieldii". Mammalian Species. 802: Number 802: pp. 1–5. doi:10.1644/802.1.
Hodgkison, R.; et al. (2004). "Habitat structure, wing morphology, and the vertical stratification of Malaysian fruit bats (Megachiroptera: Pteropodidae)". Journal of Tropical Ecology. 20 (6): 667–673. doi:10.1017/S0266467404001737. S2CID 49389740.
Campbell, P.; et al. (2006). "Comparative roosting ecology of Cynopterus (Chiroptera: Pteropodidae) fruit bats in peninsular Malaysia". Biotropica. 38 (6): 725–734. doi:10.1111/j.1744-7429.2006.00203.x. S2CID 83585694.

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