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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Cladus: Sarcopterygii
Cladus: Rhipidistia
Cladus: Tetrapodomorpha
Cladus: Eotetrapodiformes
Cladus: Elpistostegalia
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Cladus: Therapsida
Cladus: Theriodontia
Subordo: Cynodontia
Infraordo: Eucynodontia
Cladus: Probainognathia
Cladus: Prozostrodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohors: Theria
Cohors: Eutheria
Infraclassis: Placentalia
Superordo: Xenarthra
Ordo: Pilosa
Subordo: Folivora
Familia: †Scelidotheriidae
Genera: Analcitherium - Catonyx - Chubutherium - Elassotherium - Nematherium - Neonematherium - Proscelidodon - Scelidotheriops - Scelidotherium

Name
Scelidotheriidae Ameghino, 1889

Scelidotheriidae is a family of extinct ground sloths within the order Pilosa, suborder Folivora and superfamily Mylodontoidea, related to the other extinct mylodontoid family, Mylodontidae, as well as to the living two-toed sloth family Choloepodidae. The only other extant family of the suborder Folivora is the distantly related Bradypodidae. Erected as the family Scelidotheriidae by Ameghino in 1889, the taxon was demoted to a subfamily by Gaudin in 1995.[1][2] However, recent collagen sequence data indicates the group is less closely related to Mylodon and Lestodon than Choloepus is, and thus it has been elevated back to full family status by Presslee et al. (2019).[3]

Taxonomy

Together with Mylodontidae, and the two-toed sloths, the scelidotheriids form the superfamily Mylodontoidea. Chubutherium is an ancestral and very plesiomorphic member of this family and does not belong to the main group of closely related genera.
Phylogeny

The following sloth family phylogenetic tree is based on collagen and mitochondrial DNA sequence data (see Fig. 4 of Presslee et al., 2019).[3]

  Folivora  

Megalocnidae (Caribbean sloths)

Nothrotheriidae

Megatheriidae

Megalonychidae

Bradypodidae (three-toed sloths)

Megatherioidea

  Scelidotheriidae  

Scelidotherium sp.

Scelidodon sp.

Choloepodidae
  (two-toed sloths)  

C. didactylus

C. hoffmanni

  Mylodontidae  

Lestodon armatus

Paramylodon harlani

Mylodon darwinii

Glossotherium robustus    

Mylodontoidea


Below is a more detailed cladogram of the Scelidotheriidae, based on the work of Nieto et al. 2020.[4]

Scelidotheriinae 

Sibyllotherium guenguelianum

Neonematherium flabellatum

Scelidotherium

Proscelidodon gracillimus

Proscelidodon patrius

Proscelidodon rothi

Valgipes bucklandi

Catonyx cuvieri

Catonyx tarijensis

Catonyx chiliense




References

iconPaleontology portal Prehistoric mammals portal

PaleoBiology Database: Scelidotheriinae, basic info
Gaudin, T. J. (1995-09-14). "The Ear Region of Edentates and the Phylogeny of the Tardigrada (Mammalia, Xenarthra)". Journal of Vertebrate Paleontology. 15 (3): 672–705. doi:10.1080/02724634.1995.10011255. JSTOR 4523658.
Presslee, S.; Slater, G. J.; Pujos, F.; Forasiepi, A. M.; Fischer, R.; Molloy, K.; Mackie, M.; Olsen, J. V.; Kramarz, A.; Taglioretti, M.; Scaglia, F.; Lezcano, M.; Lanata, J. L.; Southon, J.; Feranec, R.; Bloch, J.; Hajduk, A.; Martin, F. M.; Gismondi, R. S.; Reguero, M.; de Muizon, C.; Greenwood, A.; Chait, B. T.; Penkman, K.; Collins, M.; MacPhee, R.D.E. (2019). "Palaeoproteomics resolves sloth relationships" (PDF). Nature Ecology & Evolution. 3 (7): 1121–1130. doi:10.1038/s41559-019-0909-z. PMID 31171860. S2CID 174813630.

Nieto, Gastón L.; Haro, J. Augusto; McDonald, H. Gregory; Miño-Boilini, Ángel R.; Tauber, Adan A.; Krapovickas, Jerónimo M.; Fabianelli, Maximiliano N.; Rosas, Federico M. (2021-06-01). "The Skeleton of the Manus of Scelidotherium (Xenarthra, Mylodontidae) Specimens from the Pleistocene of the Province of Córdoba, Argentina, and its Systematic Implications". Journal of Mammalian Evolution. 28 (2): 221–243. doi:10.1007/s10914-020-09520-x. ISSN 1573-7055.

Further reading
Cuvier, G. (1796): Notice sur le squellette d'une très grande espèce de quadrupède inconnue jusqu'à présent, trouvé au Paraquay, et déposé au cabinet d'histoire naturelle de Madrid. Magasin encyopédique, ou Journal des Sciences, des Lettres et des Arts (1): 303-310; (2): 227-228.
Iuliis, G.; Cartelle, C. (December 1999). "A new giant megatheriine ground sloth (Mammalia: Xenarthra: Megatheriidae) from the late Blancan to early Irvingtonian of Florida". Zoological Journal of the Linnean Society. 1 27 (4): 495–515. doi:10.1006/zjls.1998.0190.
Harrington, C.R. (1993): Yukon Beringia Interpretive Center - Jefferson's Ground Sloth. Retrieved 2008-JAN-24.
Hogan, C.M. (2008): Cueva del Milodon, Megalithic Portal. Retrieved 2008-APR-13
Kurtén, B.; Anderson, E. (1980). Pleistocene Mammals of North America. Columbia University Press. ISBN 978-0-231-03733-4. OCLC 776120509.
McKenna, M. C.; Bell, S. K. (1997). Classification of Mammals: Above the Species Level. Columbia University Press. ISBN 978-0-231-52853-5. OCLC 182575570.
Nowak, R.M. (1999): Walker's Mammals of the World (Vol. 2). Johns Hopkins University Press, London.
White, J. (1993). "Indicators of locomotor habits in xenarthrans: Evidence for locomotor heterogeneity among fossil sloths". Journal of Vertebrate Paleontology. 13 (2): 230–242. doi:10.1080/02724634.1993.10011502.
White, J.L.; MacPhee, R.D.E. (2001). "The sloths of the West Indies: a systematic and phylogenetic review". In Woods, C.A.; Sergile, F.E. (eds.). Biogeography of the West Indies: Patterns and Perspectives. Boca Raton, London, New York, and Washington, D.C.: CRC Press. pp. 201–235. doi:10.1201/9781420039481-14. ISBN 978-0-8493-2001-9.
Woodward, A. S. (January 1900). "On some Remains of Grypotherium (Neomylodori) listai and associated Mammals from a Cavern near Consuelo Cove, Last Hope Inlet, Patagonia". Proceedings of the Zoological Society of London. 69 (1): 64–78. doi:10.1111/j.1096-3642.1890.tb01704.x.

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