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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Ordo: Therapsida
Cladus: Theriodontia
Subordo: Cynodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohort: Theria
Cohort: Eutheria
Cohort: Placentalia
Cladus: Boreoeutheria
Superordo: Euarchontoglires
Ordo: Rodentia
Subordo: Myomorpha
Superfamilia: Muroidea

Familia: Muridae
Subfamilia: Murinae
Genus: Tokudaia
Species: Tokudaia osimensis

Tokudaia osimensis (Abe, 1933)

Type locality: Mt. Kiyako-kan, Sumiyō Village, Amami Ōshima, Kagoshima Prefecture, Japan.

Holotype: "four specimens without registration, and presumably destroyed in Hiroshima in 1945" [Kaneko, 2001: 31]

Acanthomys osimensis Tokuda, 1941: 95
Rattus fulvescens osimensis Ellerman, 1941: 193
Rattus jerdoni osimensis Abe, 1933: 942 (protonym)
Tokudaia osimensis Kuroda, 1943: 61
Tokudamys osimensis osimensis Johnson, 1946: 169

Primary references

Abe, Y. 1933. アマミトゲネズミに就いて [On the Amami Ōshima spiny rat.] Botany and Zoology 1: 936–942. Reference page.
Ellerman, J.R. 1941. The families and genera of living rodents. Volume 2. Family Muridae. Trustees of the British Museum (Natural History): London. xii + 690 pp. BHL Reference page.
Tokuda, M. A revised monograph of the Japanese and Manchou-Korean Muridae. Biogeographica: transactions of the Biogeographical Society of Japan 4(1): 1–155. Reference page.
Kuroda, N. 1943. アマミトゲネズミ沖縄島に発見せらる [The occurrence of the spinous rat in Okinawa Island in the central Riu Kiu Islands.] Bulletin of the Biogeographical Society of Japan 13: 59–64. Reference page.
Johnson, D.H. 1946. The spiny rats of the Riu Kiu Islands. Proceedings of the Biological Society of Washington 59: 169–172. BHL Reference page.

Additional references

Kaneko, Y. 2001. Morphological discrimination of the Ryukyu spiny rat (genus Tokudaia) between the islands of Okinawa and Amami Oshima, in the Ryukyu Islands, southern Japan. Mammal Study 26(1): 17–33. DOI: 10.3106/mammalstudy.26.17 Reference page.


IUCN: Tokudaia osimensis (Abe, 1933) (Endangered)

Vernacular names
Deutsch: Amami-Inselstachelratte
English: Amami Spiny Rat
español: Ratta espinosa de Amami
français: Mulot d'Amami
日本語: アマミトゲネズミ
The Ryukyu spiny rat (Tokudaia osimensis) is a species of rodent in the family Muridae.[2] Endemic to Amami Ōshima island in the Amami Islands of the Ryukyu archipelago of Japan, its natural habitat is subtropical moist broadleaf forest. The karyotype has an odd diploid number, 2n = 25.[2] Like its relative T. tokunoshimensis, it has lost its Y chromosome and SRY gene.[3]

The species is threatened by habitat destruction and fragmentation, predation by feral cats and dogs and introduced mongooses, and competition with introduced black rats.[1]
See also

Ellobius lutescens
Ellobius tancrei


Ishii, N. (2016). "Tokudaia osimensis". IUCN Red List of Threatened Species. 2016: e.T21973A22409638. doi:10.2305/IUCN.UK.2016-1.RLTS.T21973A22409638.en. Retrieved 15 November 2021.
Musser, G.G.; Carleton, M.D. (2005). "Superfamily Muroidea". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. p. 1513. ISBN 978-0-8018-8221-0. OCLC 62265494.
KAYOKO GEJI (August 23, 2012). "Y chromosome losing all genes, but men won't become extinct". The Asahi Shimbun. Archived from the original on 11 January 2014. Retrieved 11 January 2014.

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