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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Cladus: Synapsida
Cladus: Eupelycosauria
Cladus: Sphenacodontia
Cladus: Sphenacodontoidea
Ordo: Therapsida
Cladus: Theriodontia
Subordo: Cynodontia
Cladus: Mammaliaformes
Classis: Mammalia
Subclassis: Trechnotheria
Infraclassis: Zatheria
Supercohort: Theria
Cohort: Eutheria
Cohort: Placentalia
Cladus: Boreoeutheria
Superordo: Euarchontoglires
Ordo: Rodentia
Subordo: Myomorpha
Superfamilia: Muroidea

Familia: Muridae
Subfamilia: Murinae
Genus: Tokudaia
Species: Tokudaia tokunoshimensis

Tokudaia tokunoshimensis Endo & Tsuchiya, 2006: 48

Type locality: Mikyō, Amagi, Tokunoshima Island, Kagoshima Prefecture, Japan (forest).

Holotype: NSMT-M33901
Primary references

Endo, H. & Tsuchiya, K. 2006. A new species of Ryukyu spiny rat, Tokudaia (Muridae: Rodentia), from Tokunoshima Island, Kagoshima Prefecture, Japan. Mammal Study 31(1): 47–57. DOI: 10.3106/1348-6160(2006)31[47:ANSORS]2.0.CO;2 Reference page.


IUCN: Tokudaia tokunoshimensis Endo & Tsuchiya, 2006 (Endangered)

Vernacular names
Deutsch: Tokunoshima-Stachelratte
English: Tokunoshima Spiny Rat
español: Ratta espinosa de Tokunoshima
français: Mulot de Tokunoshima
日本語: トクノシマトゲネズミ

The Tokunoshima spiny rat (Tokudaia tokunoshimensis) is a rodent found only on the island of Tokunoshima in the Ryukyu Islands of Japan. Due to its small habitat, it is considered endangered. It is commonly found in the secondary and primary subtropical moist broadleaf forests of this island.[1] The karyotype has an odd diploid number, 2n = 45.[2] Like its relative T. osimensis, it is one of the few mammals that lack a Y chromosome and SRY gene.[3]

The species is threatened by deforestation and predation by feral cats and dogs.[1]
See also

Ellobius lutescens
Ellobius tancrei


Laginha Pinto Correia, D. (2016). "Tokudaia tokunoshimensis". IUCN Red List of Threatened Species. 2016: e.T136695A22409747. doi:10.2305/IUCN.UK.2016-1.RLTS.T136695A22409747.en. Retrieved 19 November 2021.
Musser, G.G.; Carleton, M.D. (2005). "Tokudaia muenninki". In Wilson, D.E.; Reeder, D.M (eds.). Mammal Species of the World: A Taxonomic and Geographic Reference (3rd ed.). Johns Hopkins University Press. pp. 1512–1513. ISBN 978-0-8018-8221-0. OCLC 62265494.
Kuroiwa A, Handa S, Nishiyama C, Chiba E, Yamada F, Abe S, Matsuda Y (2011-06-08). "Additional copies of CBX2 in the genomes of males of mammals lacking SRY, the Amami spiny rat (Tokudaia osimensis) and the Tokunoshima spiny rat (Tokudaia tokunoshimensis)". Chromosome Res. 19 (5): 635–44. doi:10.1007/s10577-011-9223-6. PMID 21656076.

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