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Gloydius tsushimaensis on Jeju Island 22

Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Superclassis: Tetrapoda
Cladus: Reptiliomorpha
Cladus: Amniota
Classis: Reptilia
Cladus: Eureptilia
Cladus: Romeriida
Subclassis: Diapsida
Cladus: Sauria
Infraclassis: Lepidosauromorpha
Superordo: Lepidosauria
Ordo: Squamata
Subordo: Serpentes
Infraordo: Caenophidia
Superfamilia: Viperoidea

Familia: Viperidae
Subfamilia: Crotalinae
Genus: Gloydius
Species: Gloydius tsushimaensis

Gloydius tsushimaensis (Isogawa, Moriya & Mitsui, 1994)

Holotype: OMNH R3934, adult ♂, collected by YMS Research team on 28 July 1987.
Type locality: “Yora-Naiin, Izuhara-cho, Shimoagata-gun, Nagasaki Prefecture, Japan”.


Agkistrodon tsushiamensis Isogawa, Moriya & Mitsui, 1994: 102 [original combination]
Gloydius tsushimaensis — Gumprecht, Tillack, Orlov, Captain & Ryabow, 2004 [subsequent combination]

Native distribution areas

Japan: Tsushima Islands.

Primary references

Isogawa, K., Moriya, A. & Mitsui, S. 1994. A new snake of the genus Agkistrodon (Serpentes, Viperidae) from Tsushima Island, Nagasaki Prefecture, Japan. Japanese Journal of Herpetology 15(30): 101–111. DOI: 10.5358/hsj1972.15.3_101 Reference page.
Gumprecht, A., Tillack, F., Orlov, N., Captain, A. & Ryabov, S. 2004. Asian Pitvipers. Geitje Books: Berlin. 368 pp. ISBN 978-3937975009 Reference page.


Uetz, P. & Hallermann, J. 2021. Gloydius tsushimaensis. The Reptile Database. Accessed on 1 May 2019.
Ota, H. & Kidera, N. 2018. IUCN: Gloydius tsushimaensis (Least Concern). The IUCN Red List of Threatened Species 2018: e.T88793478A88793486. DOI: 10.2305/IUCN.UK.2018-2.RLTS.T88793478A88793486.en

Vernacular names
English: Tsushima Island Pitviper
日本語: ツシママムシ

Gloydius tsushimaensis, or the Tsushima Island pitviper, is a species of venomous snake in the family Viperidae.[3][4] The species is endemic to Tsushima Island in Japan.[2]

Isogawa K, Moriya A, Mitsui S (1994). "A new snake from the genus Agkistrodon (Serpentes: Viperidae) from Tsushima Island, Nagasaki Prefecture". Japapanese J. Herpetol. 15: 101-111.
Gloydius tsushimaensis at the Reptarium.cz Reptile Database. Accessed 14 February 2015.
McDiarmid RW, Campbell JA, Touré T (1999). Snake Species of the World: A Taxonomic and Geographic Reference, Volume 1. Washington, District of Columbia: Herpetologists' League. 511 pp. ISBN 1-893777-00-6 (series). ISBN 1-893777-01-4 (volume).
"Gloydius ". Integrated Taxonomic Information System. Retrieved 17 October 2006.

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