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Superregnum: Eukaryota
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Superphylum: Deuterostomia
Phylum: Chordata
Cladus: Craniata
Subphylum: Vertebrata
Infraphylum: Gnathostomata
Megaclassis: Osteichthyes
Superclassis/Classis: Actinopterygii
Classis/Subclassis: Actinopteri
Subclassis/Infraclassis: Neopterygii
Infraclassis: Teleostei
Megacohors: Osteoglossocephalai
Supercohors: Clupeocephala
Cohors: Euteleosteomorpha
Subcohors: Neoteleostei
Infracohors: Eurypterygia
Sectio: Ctenosquamata
Subsectio: Acanthomorphata
Divisio/Superordo: Acanthopterygii
Subdivisio: Percomorphaceae
Series: Gobiaria
Ordo: Gobiiformes
Subordo: Gobioidei

Familia: Xenisthmidae
Genera (7): Allomicrodesmus – Gymnoxenisthmus – Kraemericus – Paraxenisthmus – Rotuma – Tyson – Xenisthmus

Xenisthmidae Miller, 1973

Miller, P.J. 1973. The osteology and adaptive features of Rhyacichthys aspro (Teleostei: Gobioidei) and the classification of gobioid fishes. Journal of zoology 171: 397–434. DOI: 10.1111/j.1469-7998.1973.tb05347.x


Xenisthmidae and its species in FishBase,
Froese, R. & Pauly, D. (eds.) 2021. FishBase. World Wide Web electronic publication, www.fishbase.org, version 02/2021.
Genera of Xenisthmidae (including synonyms) in Catalog of Fishes, Eschmeyer, W.N., Fricke, R. & van der Laan, R. (eds.) 2021. Catalog of Fishes electronic version.

Vernacular names
čeština: Pyskatcovití
日本語: ヤナギハゼ科

Collared wrigglers are perciform fishes in the family Xenisthmidae. They are native to the Indian and Pacific Oceans, where they are mostly reef-dwelling.

The 10 species in 7 genera are:

Genus Allomicrodesmus Schultz, 1966

Allomicrodesmus dorotheae Schultz, 1966 [1]

Genus Gymnoxenisthmus Gill, Bogorodsky & Mal, 2014 [2]

Gymnoxenisthmus tigrellus Gill, Bogorodsky & Mal, 2014 [2]

Genus Kraemericus

Kraemericus smithi Menon & Talwar, 1972

Genus Paraxenisthmus Gill & Hoese, 1993

Paraxenisthmus springeri Gill & Hoese, 1993 [3]

Genus Rotuma Springer, 1988

Rotuma lewisi Springer, 1988 [4]

Genus Tyson Springer, 1983

Tyson belos Springer, 1983 [5]

Genus Xenisthmus Snyder, 1908

Xenisthmus africanus Smith, 1958 (the flathead wriggler or African wriggler)[6]
Xenisthmus balius Gill & Randall, 1994 [7]
Xenisthmus chi Gill & Hoese, 2004 [8]
Xenisthmus clarus (Jordan & Seale, 1906) [9]
Xenisthmus eirospilus Gill & Hoese, 2004 [8]
Xenisthmus polyzonatus (Klunzinger, 1871) (the bull's-eye wriggler or polyzonate wriggler) is peach-coloured with a bull's-eye-like spot on its tail.
Xenisthmus semicinctus Gill & Hoese, 2004 [8]


Schultz, L.P. (1966). Schultz, L.P.; Woods, L.P.; Lachner, E.A. (eds.). Order Percomorphida. Suborder Gobiina. Superfamily Gobioidea (Print). In Fishes of the Marshall and Marianas Islands. 3. Families Kraemeridae through Antennariidae. United States: United States National Museum Bulletin. pp. 1–176.
Gill, A.C.; Bogorodsky, S.V. & Mal, A.O. (2014). "Gymnoxenisthmus tigrellus, new genus and species of gobioid fish from the Red Sea (Gobioidei: Xenisthmidae)" (PDF). Zootaxa. 3755 (5): 491–495. doi:10.11646/zootaxa.3755.5.9. PMID 24869837.
Gill, Anthony C.; Hoese, Douglass F. (1993). "Paraxenisthmus springeri, New Genus and Species of Gobioid Fish from the West Pacific, and Its Phylogenetic Position within the Xenisthmidae". Copeia. 1993 (4): 1049–1057. doi:10.2307/1447083. JSTOR 1447083.
Springer, V.G. (1988). "Rotuma lewisi, new genus and species of fish from the southwest Pacific (Gobioidei, Xenisthmidae)". Proceedings of the Biological Society of Washington. Biological Society of Washington. 101 (3): 530–539.
Springer, V.G. (1983). "Tyson belos, New Genus and Species of Western Pacific Fish (Gobiidae, Xenisthminae) : With Discussions of Gobioid Osteology and Classification" (PDF). Smithsonian Contributions to Zoology. Smithsonian Institution Press. 390 (390): 1–40. doi:10.5479/si.00810282.390. Retrieved 26 October 2012.
Smith, J.L.B. (1958). "The fishes of the family Eleotridae in the western Indian Ocean". Ichthyological Bulletin of the J.L.B. Smith Institute of Ichthyology. J.L.B. Smith Institute of Ichthyology. 11: 137–163. hdl:10962/d1018772.
Gill, Anthony C.; Randall, J.E. (1994). "Xenisthmus balius, a new species of fish from the Persian Gulf (Gobioidei: Xenisthmidae)". Proceedings of the Biological Society of Washington. Biological Society of Washington. 107 (3): 445–450.
Gill, Anthony C.; Hoese, Douglass F. (2004). "Three New Australian Species of the Fish Genus Xenisthmus (Gobioidei: Xenisthmidae)" (PDF). Records of the Australian Museum. The Australian Museum. 56 (2): 241–246. doi:10.3853/j.0067-1975.56.2004.1428. ISSN 0067-1975. Retrieved 26 October 2012.

Jordan, D.S.; Seale, A. (1994). "he fishes of Samoa. Description of the species found in the archipelago, with a provisional check-list of the fishes of Oceania". Bulletin of the Bureau of Fisheries. 25.

Froese, Rainer, and Daniel Pauly, eds. (2006). "Xenisthmidae" in FishBase. January 2006 version.

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