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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Heterobranchia
Infraclassis: Euthyneura
Cohors: Tectipleura
Subcohors: Panpulmonata
Superordo: Eupulmonata
Ordo: Stylommatophora
Subordo: Helicina
Infraordo: Limacoidei
Superfamilia: Helicarionoidea

Familia: Ariophantidae
Subfamiliae (3): Ariophantinae – Macrochlamydinae – Ostracolethinae

Genus Incertae sedis (1): Falsiplecta
Overview of genera (50)

Apoparmarion – Ariophanta – Baiaplecta – Bapuia – Cambodiparmarion – Cryptogirasia – Cryptosemelus – Cryptozona – Dalingia – Damayantia – Damayantiella – Dekhania – Dihangia – Euplecta – Falsiplecta – Galongia – Hemiplecta – Himalodiscus – Indrella – Isselentia – Khasiella – Koratia – Macrochlamys – Mariaella – Megaustenia – Microcystina – Microparmarion – Minyongia – Muangnua – Ostracolethe – Oxytesta – Paraparmarion – Parmarion – Parmunculus – Parvatella – Philippinella – Rahula – Ratnadvipia – Ravana – Sakiella – Sarika – Schwammeria – Sesara – Sitalinopsis – Syama – Tadunia – Taphrospira – Vitrinula – Wiegmannia – Xesta
Name

Ariophantidae Godwin-Austen, 1883
References
Links

Ariophantidae in the World Register of Marine Species

Ariophantidae is a taxonomic family of air-breathing land snails and semi-slugs, terrestrial pulmonate gastropod mollusks in the superfamily Helicarionoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[2]
Distribution

The distribution of the family Ariophantidae includes India and south-eastern Asia.[3]
Anatomy

Snails within this family make and use love darts made of chitin.[4]

In this family, the number of haploid chromosomes lies between 21 and 25 and also lies between 31 and 35, but other values are also possible (according to the values in this table).[5]
Taxonomy
Subfamilies

The family Ariophantidae consists of 3 subfamilies:

Ariophantinae Godwin-Austen, 1888 - synonyms: Naninidae Pfeffer, 1878 (inv.); Hemiplectinae Gude & B. B. Woodward, 1921[6]
Macrochlamydinae Godwin-Austen, 1888[1] - synonyms: Tanychlamydinae H. B. Baker, 1928;[7] Vitrinulini Schileyko, 2003[8]
Ostracolethinae Simroth, 1901 - synonyms: Parmarioninae Godwin-Austen, 1908;[9] Laocaiini Schileyko, 2002;[10] Microparmarionini Schileyko, 2003;[11] Myotestidae Collinge, 1902[12]

Genera

The following genera are recognised in the family Ariophantidae:[13]

Ariophantinae

Ariophanta Des Moulins, 1829 - type genus of the family Ariophantidae[2]
Bapuia Godwin-Austen, 1918
Cryptogirasia Cockerell, 1891
Cryptozona Mörch, 1872
Dalingia Godwin-Austen, 1907
Dihangia Godwin-Austen, 1916
Euplecta C. Semper, 1870
Galongia Godwin-Austen, 1916
Hemiplecta Albers, 1850
Indrella Godwin-Austen, 1901
Khasiella Godwin-Austen, 1899
Mariaella Gray, 1855
Megaustenia Cockerell, 1912
Ratnadvipia Godwin-Austen, 1899 - endemic to Sri Lanka[14]
Ravana Godwin-Austen, 1901
Schwammeria Schileyko, 2010
Sitalinopsis Thiele, 1931
Taphrospira W. T. Blanford, 1904
Xesta Albers, 1850

Macrochlamydinae

Baiaplecta Laidlaw, 1956
Himalodiscus Kuznetsov, 1996
Macrochlamys Gray, 1847 - type genus of the subfamily Macrochlamydinae[2]
Microcystina Mörch, 1872
Oxytesta Zilch, 1956
Parvatella W. T. Blanford & Godwin-Austen, 1908
Rahula Godwin-Austen, 1907
Sakiella Godwin-Austen, 1908
Sarika Godwin-Austen, 1907
Sesara Albers, 1860
Syama W. T. Blanford & Godwin-Austen, 1908
Tadunia Godwin-Austen, 1918
Taphrenalla Pholyotha & Panha, 2021
Varadia Bhosale & Raheem, 2021
Vitrinula Gray, 1857

Microparmarion simrothi

Ostracolethinae

Apoparmarion Collinge, 1902
Cambodiparmarion Kuznetsov & Kuzminykh, 1999
Cryptosemelus Collinge, 1902
Damayantia Issel, 1874
Damayantiella Zilch, 1959
Isselentia Collinge, 1901
Microparmarion Simroth, 1893
Minyongia Godwin-Austen, 1916
Ostracolethe Simroth, 1901 - type genus of the subfamily Ostracolethinae
Paraparmarion Collinge, 1902
Parmarion P. Fischer, 1855
Parmunculus Collinge, 1899
Philippinella Möllendorff, 1899
Wiegmannia Collinge, 1901

Subfamily not assigned

Falsiplecta Schileyko & Semenyuk, 2018

Cladogram

The following cladogram shows the phylogenic relationships of this family with other families in the limacoid clade:[3]

 limacoid clade 
 Staffordioidea 

Staffordiidae

 Dyakioidea 

Dyakiidae

 Gastrodontoidea 

Pristilomatidae

Chronidae

Euconulidae

Trochomorphidae

Gastrodontidae

Oxychilidae

 Parmacelloidea 

Trigonochlamydidae

Parmacellidae

Milacidae

 Zonitoidea 

Zonitidae

 Helicarionoidea 

Helicarionidae

Ariophantidae

Urocyclidae

 Limacoidea 

Vitrinidae

Boettgerillidae

Limacidae

Agriolimacidae

References

Godwin-Austen H. H. (April 1888). Land and freshwater Mollusca of India. including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean, supplementary to Messrs. Theobald and Hanley's Conchologia Indica. Taylor & Francis, London, volume 1, part 6: 207-257, page 253.
Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
Hausdorf B. (2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.
"Bringing the Lab to the Field: A New Lowland Microparmarion Semi-slug (Gastropoda: Ariophantidae) from Borneo, described and DNA-barcoded in the Forest". Novataxa. Retrieved 11 February 2019.
Barker G. M. (2001). Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.
Gude G. P. L. K. & Woodward B. B. (1921). "On Helicella, Férussac". Proceedings of the Malacological Society of London 14(5-6): 174-190. page 186.
Baker H. B. (1928). Proceedings of the Academy of Natural Sciences of Philadelphia 80: 6.
Schileyko (2003). Treatise on Recent terrestrial pulmonate molluscs, part 10: page 1354
Godwin-Austen H. H. (1908). In: Blanford W. T. & Godwin-Austen H. H. (1908). The fauna of British India. Mollusca. Testacellidae and Zonitidae, page 180.
Schileyko (2002). Treatise on Recent terrestrial pulmonate molluscs, part 9: page 1219.
Schileyko (2003). Treatise on Recent terrestrial pulmonate molluscs, part 10: page 1337.
"WoRMS - World Register of Marine Species - Ostracolethinae Simroth, 1901". www.marinespecies.org. Retrieved 2022-03-27.
"WoRMS - World Register of Marine Species - Ariophantidae Godwin-Austen, 1883". www.marinespecies.org. Retrieved 2022-03-27.

Raheem D. & Naggs F. (2006). "The Sri Lankan endemic semi-slug Ratnadvipia (Limacoidea: Ariophantidae) and a new species from southwestern Sri Lanka". Systematics and Biodiversity 4(1): 99-126. doi:10.1017/S1477200005001854.

Further reading

Schileyko A. A. (2003). "Treatise on recent terrestrial pulmonate mollusks. 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae". Ruthenica, Supplement 2. 1309-1466.

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