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Cladus: Eukaryota
Supergroup: Opisthokonta
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropodae
Subclassis: Orthogastropoda
Superordo: Heterobranchia
Ordo: Pulmonata
Subordo: Eupulmonata
Clade: Stylommatophora
Cladus: Orthurethra
Informal group: Sigmurethra
Superfamilia: Dyakioidea
Familia: Dyakiidae
Subfamilia: Dyakiinae ...


Dyakiidae Gude & Woodward, 1921


Dyakiidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Dyakioidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[2]

Dyakiidae is the only family in the superfamily Dyakioidea.[2] This family has no subfamilies.[2]


The family Dyakiidae is endemic to Sundaland in Southeast Asia.[3]

In this family, the number of haploid chromosomes lies between 26 and 30 (according to the values in this table).[4]

The family also include Quantula striata, the only known terrestrial gastropod to emit light.[5]

Digestive system characteristics: Buccal mass is small. The jaw is smooth. The stomach is very simple with weak muscles (as is the case in the majority of land snails).[4] (This anatomy characteristics include family Staffordiidae as as a part of Dyakiidae).

The family Dyakiidae includes the following 11 genera, with altogether 82 species (71 species up to 2007[6] + 11 new species in 2009[7]):

Asperitas Gray, 1857 - 14 species
Bertia Ancey, 1887 - only one species: Bertia cambojiensis (Reeve, 1860)
Dyakia Godwin-Austen, 1891 - type genus, 22 species
Elaphroconcha Gude, 1911 - 10 species
Everettia Godwin-Austen, 1891[8] - 15 species
Kalamantania Laidlaw, 1931 - only one species: Kalamantania whiteheadi (Godwin-Austen, 1891)[9]
Phuphania Tumpeesuwan, Naggs & Panha, 2007 - only one species: Phuphania globosa Tumpeesuwan, Naggs & Panha, 2007[6]
Pseudoplecta Laidlaw, 1932 - only one species: Pseudoplecta bijuga (Stoliczka, 1873)
Quantula Baker, 1941 - only one species: Quantula striata (Gray, 1834)
Rhinocochlis Thiele, 1931 - only one species: Rhinocochlis nasuta (Metcalfe, 1852)
Sasakina Rensch, 1930 - four species


The following cladogram shows the phylogenic relationships of this family and superfamily with the othe

 limacoid clade 



























^ a b Gude G. P. L. K. & Woodward B. B. (1921). "On Helicella, Férussac". Proceedings of the Malacological Society of London 14(5-6): 174-190. page 185.
^ a b c Bouchet P., Rocroi J.-P., Frýda J., Hausdorf B., Ponder W., Valdés Á. & Warén A. (2005). "Classification and nomenclator of gastropod families". Malacologia: International Journal of Malacology (Hackenheim, Germany: ConchBooks) 47 (1-2): 1–397. ISBN 3925919724. ISSN 0076-2997.
^ Hausdorf D. (1995). "A preliminary phylogenetic and biogeographic analysis of the Dyakiidae (Gastropoda: Stylommatophora) and a biogeographic analysis of other Sundaland taxa". Cladistics 11(4): 359 - 376. doi:10.1111/j.1096-0031.1995.tb00095.x.
^ a b Barker G. M. (2001) Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. 1-146. In: Barker G. M. (ed.) (2001) The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, cited pages: 139-144. ISBN 0-85199-318-4.
^ Isobe M., Uyakul D., Goto T. & Counsilman J. J. (April/June 1988) "Dyakia bioluminescence - 1. Bioluminescence and fluorescence spectra of the land snail, D. striata". Journal of Bioluminescence and Chemiluminescence 2(2): 73 - 79. doi:10.1002/bio.1170020204, PubMed.
^ a b Tumpeesuwan C., Naggs F. & Panha S. (31 August 2007) "A new genus and new species of dyakiid snail (Pulmonata: Dyakiidae) from the Phu Phan range, northeastern Thailand". Raffles Bulletin of Zoology 55(2): 363-369. PDF
^ Liew T.-S., Schilthuizen M. & Vermeulen J. J. (November 2009, online: 26 October 2009) "Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo". Zoological Journal of the Linnean Society 157(3): 515-550. doi:10.1111/j.1096-3642.2009.00526.x.
^ Godwin-Austen H. H. (1891) "On a collection of Land-shells made In Borneo by Mr. A. Everett, with Descriptions of supposed new species. Part II. Zonitidae and Helicidae". Proceedings of the Zoological Society of London for the year 1891: 22-47, p. 33.
^ Godwin-Austen H. H. (1891) "On a collection of Land-shells made In Borneo by Mr. A. Everett, with Descriptions of supposed new species. Part II. Zonitidae and Helicidae". Proceedings of the Zoological Society of London for the year 1891: 22-47, p. 24.
^ Hausdorf B. (March 2000). "Biogeography of the Limacoidea sensu lato (Gastropoda: Stylommatophora): Vicariance Events and Long-Distance Dispersal". Journal of Biogeography 27(2): 379-390. doi:10.1046/j.1365-2699.2000.00403.x, JSTOR.

Further reading

Laidlaw F. F. (1931). "On a new sub-family Dyakiinae of the Zonitidae". Proceedings of the malacological Society of London 19: 190-201. abstract.
Schileyko A. A. (2003). "Treatise on recent terrestrial pulmonate mollusks. 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae". Ruthenica, Supplement 2. 1309-1466.

External links

Dyakiidae at National Center for Biotechnology Information (NCBI) website.

Biology Encyclopedia

Mollusca Images

Source: Wikipedia, Wikispecies: All text is available under the terms of the GNU Free Documentation License