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Superregnum: Eukaryota
Cladus: Unikonta
Cladus: Opisthokonta
Cladus: Holozoa
Regnum: Animalia
Subregnum: Eumetazoa
Cladus: Bilateria
Cladus: Nephrozoa
Cladus: Protostomia
Cladus: Spiralia
Cladus: Lophotrochozoa
Phylum: Mollusca
Classis: Gastropoda
Subclassis: Heterobranchia
Infraclassis: Euthyneura
Cohors: Tectipleura
Subcohors: Panpulmonata
Superordo: Eupulmonata
Ordo: Stylommatophora
Subordo: Helicina
Superfamilia Incertae sedis: Urocoptoidea

Familia: Urocoptidae
Subfamiliae (2): BrachypodellinaeUrocoptinae

Genus Incertae sedis (1): Cylindrella
Overview of genera (68)

Acrocoptis – Allocoptis – Amphicosmia – Amphistemma – Anafecoptis – Angulicervix – Anoma – Apoma – Arangia – Archegocoptis – Autocoptis – Bactrocoptis – Badiofaux – Bialasmus – Braychpodella – Brevipedella – Callocoptis – Callonia – Capillacea – Carcinostemma – Centralia – Cochlodinella – Cyclauchen – Cylindrella – Esochara – Fibricutis – Geminicoptis – Geoscala – Gongylostoma – Gongylostomella – Gyraxis – Heterocoptis – Idiostemma – Insulaceramus – Ischnostrophina – Johaniceramus – Levistemma – Liocallonia – Liparotes – Macroceramus – Microceramus – Mychostoma – Nesocoptis – Nodulia – Organocoptis – Orienticoptis – Paracallonia – Pfeiffericoptis – Pineria – Planostemma – Pleurostemma – Poecilocoptis – Poeycoptis – Pycnoptychia – Sagracoptis – Sectilumen – Simplicervix – Spiroceramus – Steatocoptis – Strophina – Teneria – Tenuistemma – Tetrentodon – Tomelasmus – Torrecoptis – Trilamellaxis – Uncinicoptis – Urocoptis
Name

Urocoptidae Pilsbry, 1898 (1868)
References
Additional references

Kabat, A.R.; Hershler, R.; González-Guillén, A. 2012: Resolution of taxonomic problems associated with the complex publication history of the seminal Torre and Bartsch monograph on Cuban Urocoptidae (Gastropoda, Pulmonata). Zootaxa 3362: 43–53. Preview PDF Reference page.
Thompson, F.G. & Correa-Sandoval, A. 2011. Another new species of the urocoptid land snail genus Hendersoniella (Pulmonata, Urocoptidae, Holospirinae) from northeastern Mexico. Veliger 51(2): 91–95.

Links

Urocoptidae in the World Register of Marine Species

Urocoptidae is a family of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Urocoptoidea.[2]
Contents

1 Taxonomy
1.1 2005 taxonomy
1.2 2008 taxonomy
1.3 2012 taxonomy
2 Genera
3 References
4 Further reading
5 External links

Taxonomy
2005 taxonomy

The family Urocoptidae was classified in the superfamily Orthalicoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[3]

The family Urocoptidae consists of 7 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005):

Urocoptinae Pilsbry, 1898 (1868) - synonym: Cylindrellidae Tryon, 1868
Apomatinae Paul, 1982
Brachypodellinae H. B. Baker, 1956
Eucalodiinae P. Fischer & Crosse, 1873
Holospirinae Pilsbry, 1946
Microceraminae Pilsbry, 1904 - synonyms: Johaniceraminae Jaume & de la Torre, 1972; Macroceraminae Jaume & de la Torre, 1972
Tetrentodontinae Bartsch, 1943

2008 taxonomy

Uit de Weerd (2008)[2] moved the Urocoptidae to the newly established superfamily Urocoptoidea based on molecular phylogeny research.[2]

The following cladogram based on Bayesian analysis of sequences of 28S ribosomal RNA shows the phylogenic relations within the Urocoptoidea.[2] There are up to 2008 genetically analyzed species of Urocoptidae and one species of Cerionidae marked in khaki color in the cladogram.[2]

Urocoptoidea

Macroceramus microdon

Microceramus pontificus

Torrecoptis holguinensis

Autocoptis menkeana

Simplicervix inornata

Brachypodella dominicensis gabbi

Dissotropis sp. nov. [1]

Anisospira lebmanni

Archegocoptis crenata

Eucalodium sp.

Cerionidae

Cerion striatellum

Coelostemma sp. nov. [2]

Hendersoniella lux lux

Holospira fortisculpta

Epirobia polygyra



2012 taxonomy

Fred G. Thompson established the family Epirobiidae in 2012 for five genera of snails (Epirobia, Propilsbrya, Pectinistemma, Gyrocion, Prionoloplax), that were previously placed within Urocoptidae.[4] Data from molecular phylogeny research as well as morphology supports monophyly of Holospiridae Pilsbry, 1946 and the Eucalodiidae Crosse & Fischer, 1868.[4]
Genera

Genera in the family Urocoptidae include:

Urocoptinae

Autocoptis Pilsbry, 1902[2]
Brachypodella L. Pfeiffer, 1840 (as Cylindrella)
Urocoptis Beck, 1837 - type genus of the family Urocoptidae[3]

Apominae

Apoma Beck, 1837 - type genus of the subfamily Apomatinae[3]
Simplicervix Pilsbry, 1903[2]

Brachypodellinae

Brachypodella Beck, 1837 - type genus of the subfamily Brachypodellinae[3]

Eucalodiinae

Anisospira Strebel, 1880[2]
Archegocoptis Pilsbry, 1903[2]
†Condonella McLellan, 1927[5]
Dissotropis Bartsch, 1906[2]
Eucalodium Crosse & P. Fischer, 1868 - type genus of the subfamily Eucalodiinae[3]

Holospirinae

Coelostemma Dall, 1895[2]
Hendersoniella Dall, 1905[2]
Holospira Martens, 1860 - type genus of the subfamily Holospirinae[3]

Microceraminae

Microceramus Pilsbry & Vanatta, 1898 - type genus of the subfamily Microceraminae[3]
Macroceramus Guilding, 1828[2]

Tetrentodontinae

Tetrentodon Pilsbry, 1903 - type genus of the subfamily Tetrentodontinae[3]
Torrecoptis Bartsch, 1943[2]

Other genera include:[6]

Allocoptis Thompson & Franz, 1976
Amphicosmia Pilsbry & Vanatta, 1898
Anoma
Arangia
Bactrocoptis
Badiofax
Bostrichocentrum
Bostrychocentrum
Callonia
Centralia
Cochlodinella
Coelocentrum
Geoscala
Gongylostoma
Gyraxis
Haplocion
Heterocoptis
Idiostemma
Liocallonia
Metastoma
Mychostoma
Nesocoptis
Nodulia
Oligostylus
Organocoptis
Pfeiffericoptis
Pineria
Pleurostemma
Poeycoptis
Pseudopineria
Pycnoptychia Pilsbry & Vanatta, 1898
Sagracoptis
Spiroceramus
Spirostemma
Strophina
Tomelasmus
Trilamellaxis
Uncinicoptis

References

Pilsbry H. A. (1898). The Nautilus 11(9): 107.
Uit De Weerd, D. R. (2008). "Delimitation and phylogenetics of the diverse land-snail family Urocoptidae (Gastropoda: Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion". Journal of Molluscan Studies. 74 (4): 317–29. doi:10.1093/mollus/eyn023.
Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. Hackenheim, Germany: ConchBooks. 47 (1–2): 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
Thompson F. G. (2012). "The land snail genus Epirobia and allied genera in México and Central America, with the description of a new family, the Epirobiidae (Gastropoda, Pulmonata, Urocoptoidea)". Bulletin Florida Museum of Natural History 51: 167-215. PDF.
Roth, B. (2000). "Upper Cretaceous (Campanian) land snails (Gastropoda: Stylommatophora) from Washington and California". Journal of Molluscan Studies. 66 (3): 373–381. doi:10.1093/mollus/66.3.373.

http://pick4.pick.uga.edu/mp/20q?search=Urocoptidae

Further reading

Bartsch, P. (1943). "Notes on Mexican Urocoptid mollusks". Journal of the Washington Academy of Sciences. 33 (2): 54–59.; 7 figs.
Carlos de la Torre. 1912. New Cuban Urocoptis of the U. cinerea group. The Nautilus, volume 26, number 4, pages 55-58, plate 6.
Kabat A. R., Herschler R. & González G. A. (2012). "Resolution of taxonomic problems associated with the complex publication history of the seminal Torre and Bartsch monograph on Cuban Urocoptidae (Gastropoda, Pulmonata)". Zootaxa 3362: 43-53. PDF.
Pilsbry H. A. & Henderson J. B. jr. (February 1913). Two new Cuban Urocoptidae. The Nautilus, volume 26, number 10, pages 109-110.
(in Spanish) Miguel L. Jaume & Alfredo de la Torre. 1976. Los Urocoptidae de Cuba (Mollusca-Pulmonata). Série 4: Ciencias biologicas; no. 53. La Habana, Cuba: Centro de Información Cientifica y Tecnica, 122 pp.
Mesa Garcia, R; Jaume, ML (1982). "Zoogeography, ecology, and protection of the Urocoptidae family in Cuba (Gastropoda-Pulmonata). II". Revista Cubana de Medicina Tropical. 34 (2): 172–92. PMID 6763741.
Richardson C. L. 1991. Urocoptidae: catalogue of species. Tryonia 22: 1-245.
Tryon G. W. (1903) Manual of Conchology. Second series: Pulmonata. Volume 15. Urocoptidae. Continued by H. A. Pilsbry.

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