ART

Superregnum: Eukaryota
Regnum: Fungi
Subregnum: Dikarya
Divisio: Ascomycota
Subdivisio: Pezizomycotina
Classis: Eurotiomycetes
Subclassis: Chaetothyriomycetidae
Ordo: Verrucariales

Familia: Verrucariaceae
Genus: Atla
Species: A. alaskana – A. alpina – A. oulankaensis – A. palicei – A. praetermissa – A. recondita – A. tibelliorum – A. vitikainenii – A. wheldonii
Name

Atla Savić & Tibell, 2008


Type species: Atla alpina Savić & Tibell, 2008
References

Savić & Tibell, Lichenologist 40(4): 273 (2008)

Savić, S. and Tibell, L. (2008) Atla, a new genus in the Verrucariaceae (Verrucariales). The Lichenologist 40(4): 269–282. **[RLL List # 212 / Rec.# 30304] - (Recent Literature on Lichens)** (doi: 10.1017/S0024282908007512)

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Vernacular names

Atla is a genus of crustose lichens in the family Verrucariaceae.[1] It has nine species that grow on rocks or on soil.
Taxonomy

The genus was circumscribed in 2008 by Sanja Savić and Leif Tibell, with Atla alpina assigned as the type species. According to the authors, the generic name Atla refers to "the name of a Norse water goddess who sometimes accompanied us in the same habitats as the species named after her." Three species were included in the original circumscription of the genus; these species formed a well-defined clade in the family Verrucariaceae, sister to a clade containing several Sporodictyon species.[2] Several newly described species were later added from Alaska and Finland.[3][4]
Description

Atla lichens have crustose thalli that are blackish in colour with a surface texture ranging from warty (verrucose) to diffusely areolate to immersed in the substrate. The perithecia are 0.3–0.9 mm in diameter, and either sessile or immersed. Ascospores are ellipsoid, hyaline to dark brown when mature, and measure 43–87 by 19–49 μm; the spores are muriform (divided into compartments), with 7–15 transverse and 2–5 longitudinal walls. The excipulum (the ring-shaped layer surrounding the hymenium) is spherical, ranging in colour from black to dark brown. The involucrellum (the upper, often exposed covering or cap external to the excipulum) is well developed; depending on the species, the upper part of this structure is either fused together with the excipulum, or missing altogether.[2]
Species

Atla alaskana S.Tibell & Tibell (2015)[3] – Alaska
Atla alpina Savić & Tibell (2008)[2] – Europe
Atla oulankaensis Pykälä & Myllys (2016)[4] – Finland
Atla palicei Savić & Tibell (2008)[2] – Sweden
Atla praetermissa Savić & Tibell (2008)[2] – Scandinavia
Atla recondita S.Tibell & Tibell (2015)[3] – Sweden
Atla tibelliorum Pykälä & Myllys (2016)[4] – Finland
Atla vitikainenii Pykälä & Myllys (2016)[4] – Finland
Atla wheldonii (Travis) Savić & Tibell (2008)[2] – Europe

References

Wijayawardene, N.N.; Hyde, K.D.; Dai, D.Q.; Sánchez-García, M.; Goto, B.T.; Saxena, R.K.; et al. (2022). "Outline of Fungi and fungus-like taxa – 2021". Mycosphere. 13 (1): 53–453. doi:10.5943/mycosphere/13/1/2. S2CID 249054641.
Savić, Sanja; Tibell, Leif (2008). "Atla, a new genus in the Verrucariaceae ( Verrucariales)". The Lichenologist. 40 (4): 269–282. doi:10.1017/s0024282908007512. S2CID 85405431.
Tibell, Sanja; Tibell, Leif (2015). "Two new species of Atla (Verrucariaceae)". The Lichenologist. 47 (2): 93–98. doi:10.1017/s0024282915000018. S2CID 83611538.
Pykälä, Juha; Myllys, Leena (2016). "Three new species of Atla from calcareous rocks (Verrucariaceae, lichenized Ascomycota)". The Lichenologist. 48 (2): 111–120. doi:10.1017/s0024282915000523. S2CID 89862994.

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