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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Ordo: Ericales

Familia: Ericaceae
Subfamilia: Arbutoideae
Genera: ArbutusArctostaphylosArctousComarostaphylisOrnithostaphylosXylococcus

Name

Arbutoideae (Meisn.) Nied.
Links

International Plant Names Index. 2020. Arbutoideae. Published online. Accessed: Apr 17 2020.
Tropicos.org 2020. Arbutoideae. Missouri Botanical Garden. Published online. Accessed: 17 Apr 2020.

Vernacular names
한국어: 아르부투스아과

The Arbutoideae are a subfamily in the flowering plant family Ericaceae. Phylogenetic analysis supported all genera of the subfamily as monophyletic, except Arbutus.[1] Moreover, it was suggested that the non-sister relationship between Mediterranean and North American species may be explained by a once widespread distribution in the Northern hemisphere before the Neogene.[1]

The genera Arbutus, Arctostaphylos, Comarostaphylis form a particular type of mycorrhizal symbiosis with the fungus, Arbutoid mycorrhiza, which resembles ectomycorrhizas.[2][3]
Genera List

Image Genus Living Species
Arbutus unedo (frutu).jpg Arbutus L.
  • Arbutus andrachne – Greek strawberry tree
  • Arbutus canariensis – Canary madrone
  • Arbutus pavarii
  • Arbutus unedo – Strawberry tree
  • Arbutus arizonica – Arizona madrone
  • Arbutus bicolor
  • Arbutus madrensis
  • Arbutus menziesii – Pacific madrone
  • Arbutus mollis
  • Arbutus occidentalis
  • Arbutus tessellata
  • Arbutus xalapensis – Texas madrone
Arctostaphylos pringlei ssp drupacea 3.jpg Arctostaphylos Adans.
  • Subgenus Micrococcus
    • Sect. Micrococcus
      • Arctostaphylos mendocinoensis Pygmy manzanita
      • Arctostaphylos myrtifolia Ione manzanita
      • Arctostaphylos nissenana Nissenan manzanita
      • Arctostaphylos nummularia Glossyleaf manzanita
  • Subgenus Arctostaphylos, which has 3 sections:
    • Sect. Arctostaphylos
      • Arctostaphylos alpina Alpine bearberry
      • Arctostaphylos bakeri Baker's manzanita
      • Arctostaphylos densiflora Sonoma manzanita
      • Arctostaphylos edmundsii Little Sur manzanita
      • Arctostaphylos franciscana Franciscan manzanita
      • Arctostaphylos gabrielensis San Gabriel manzanita
      • Arctostaphylos glauca Bigberry manzanita
      • Arctostaphylos hispidula Gasquet manzanita
      • Arctostaphylos hookeri Hooker's manzanita
      • Arctostaphylos insularis Island manzanita
      • Arctostaphylos klamathensis Klamath manzanita
      • Arctostaphylos manzanita Common manzanita
      • Arctostaphylos mewukka Indian manzanita
      • Arctostaphylos nevadensis Pinemat manzanita
      • Arctostaphylos parryana Parry manzanita
      • Arctostaphylos patula Greenleaf manzanita
      • Arctostaphylos pumila Sandmat manzanita
      • Arctostaphylos pungens Pointleaf manzanita
      • Arctostaphylos rudis Shagbark manzanita
      • Arctostaphylos stanfordiana Stanford's manzanita
      • Arctostaphylos uva-ursi Bearberry
      • Arctostaphylos viscida Sticky manzanita
    • Sect. Foliobracteata
      • Arctostaphylos andersonii Santa Cruz manzanita
      • Arctostaphylos auriculata Mount Diablo manzanita
      • Arctostaphylos canescens Hoary manzanita
      • Arctostaphylos catalinae Santa Catalina Island manzanita
      • Arctostaphylos columbiana Hairy manzanita
      • Arctostaphylos confertiflora Santa Rosa Island manzanita
      • Arctostaphylos cruzensis La Cruz manzanita
      • Arctostaphylos glandulosa Eastwood manzanita
      • Arctostaphylos glutinosa Schreiber's manzanita
      • Arctostaphylos hooveri Hoover's manzanita
      • Arctostaphylos imbricata San Bruno Mountain manzanita
      • Arctostaphylos luciana Santa Lucia manzanita
      • Arctostaphylos malloryi Mallory's manzanita
      • Arctostaphylos montaraensis Montara manzanita
      • Arctostaphylos montereyensis Monterey manzanita
      • Arctostaphylos morroensis Morro manzanita
      • Arctostaphylos nortensis Del Norte manzanita
      • Arctostaphylos obispoensis Serpentine manzanita
      • Arctostaphylos osoensis Oso manzanita
      • Arctostaphylos otayensis Otay manzanita
      • Arctostaphylos pajaroensis Pajaro manzanita
      • Arctostaphylos pallida Pallid manzanita
      • Arctostaphylos pechoensis Pecho manzanita
      • Arctostaphylos pilosula La Panzo manzanita
      • Arctostaphylos purissima La Purissima manzanita
      • Arctostaphylos refugioensis Refugio manzanita
      • Arctostaphylos regismontana Kings Mountain manzanita
      • Arctostaphylos silvicola Bonny Doon manzanita
      • Arctostaphylos tomentosa Woolyleaf manzanita
      • Arctostaphylos virgata Bolinas manzanita
      • Arctostaphylos viridissima Whitehair manzanita
      • Arctostaphylos wellsii Wells' manzanita
    • Sect. Pictobracteata
      • Arctostaphylos pringlei Pringle manzanita
  • Unassigned
    • Arctostaphylos rainbowensis Rainbow manzanita
    • Arctostaphylos gabilanensis Gabilan manzanita
    • Arctostaphylos ohloneana Ohlone manzanita
Arctostaphylos alpinus (Alpen-Bärentraube) IMG 6699.JPG Arctous (A.Gray) Nied.
  • Arctous alpina (L.) Nied.
  • Arctous microphyllus C.Y.Wu
  • Arctous ruber (Rehder & E.H.Wilson) Nakai
Comarostaphylis diversifolia (4986961909).jpg Comarostaphylis Zucc.
  • Comarostaphylis arbutoides
  • Comarostaphylis discolor
  • Comarostaphylis diversifolia
  • Comarostaphylis glaucescens
  • Comarostaphylis lanata
  • Comarostaphylis longifolia
  • Comarostaphylis mucronata
  • Comarostaphylis polifolia
  • Comarostaphylis sharpii
  • Comarostaphylis spinulosa
P20150305-0094—Ornithostaphylos oppositifolia—RPBG (16550855467).jpg Ornithostaphylos Small
  • Ornithostaphylos oppositifolia, the Baja birdbush
MissionManzanitaBlooming03-P.jpg Xylococcus Nutt.
  • Xylococcus bicolor, the mission manzanita


References

Hileman LC, Vasey MC, Parker VT (2001). "Phylogeny and biogeography of the Arbutoideae (Ericaceae): implications for the Madrean–Tethyan hypothesis". Systematic Botany. 26 (1): 131–143. doi:10.1043/0363-6445-26.1.131 (inactive 28 February 2022).
Osmundsen TW, Halling RE, den Bakker HC (2007). "Morphological and molecular evidence supporting an arbutoid mycorrhizal relationship in the Costa Rican páramo". Fungal Diversity. 17 (3): 217–222. doi:10.1007/s00572-006-0098-x. PMID 17216498. S2CID 19084561.
Smith SE, Read D (2008). Mycorrhizal Symbiosis (3rd ed.). Amsterdam; Boston: Academic Press. ISBN 978-0-12-370526-6.

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