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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Lamiales

Familia: Acanthaceae
Subfamilia: Acanthoideae
Tribus: Acantheae
Subtribus: Acanthinae
Genus: Blepharis
Species: B. ablephara – B. abyssinica – B. acanthodioides – B. acuminata – B. aequisepala – B. affinis – B. angusta – B. arborea – B. aspera – B. asperrima – B. asteracantha – B. attenuata – B. baguirmiensis – B. bainesii – B. bequaertii – B. boranensis – B. bossii – B. breijeri – B. breviciliata – B. breyeri – B. buchneri – B. burundiensis – B. calaminthaefolia – B. calcitrapa – B. caloneura – B. capensis – B. carduacea – B. carduifolia – B. cataractae – B. chrysotricha – B. ciliaris – B. crinita – B. cristata – B. cuanzensis – B. cuspidata – B. decussata – B. dhofarensis – B. dichotoma – B. dilatata – B. diplodonta – B. diversispina – B. drummondii – B. dunensis – B. duvigneaudii – B. ecklonii – B. edulis – B. espinosa – B. evansii – B. exigua – B. extenuata – B. fenestralis – B. ferox – B. flava – B. fleckii – B. forgiarinii – B. frutescens – B. fruticulosa – B. furcata – B. gazensis – B. gerlindae – B. gigantea – B. glauca – B. glinus – B. glomerans – B. glomerata – B. glumacea – B. grandis – B. grisea – B. grossa – B. guiinzii – B. hildebrandtii – B. hirsuta – B. hirta – B. hirtella – B. hirtinervia – B. homblei – B. hornbyae – B. huillensis – B. ilicifolia – B. ilicina – B. inaequalis – B. inermis – B. inflata – B. innocua – B. inopinata – B. integrifolia – B. involucrata – B. itigiensis – B. javanica – B. kaessneri – B. katangensis – B. kenyensis – B. kuriensis – B. laevifolia – B. lawsonii – B. leedertaziae – B. leendertziae – B. leptophylla – B. lineariifolia – B. longifolia – B. longispica – B. macra – B. maculata – B. madagascariensis – B. madandensis – B. maderaspatensis – B. malangensis – B. marginata – B. menocotyle – B. meyeri – B. mitrata – B. molluginifolia – B. montana – B. myresii – B. naegelsbachii – B. natalensis – B. noli-me-tangere – B. obermeyerae – B. obmitrata – B. obovata – B. obtusisepala – B. ogadenensis – B. panduriformis – B. paradoxa – B. pascuorum – B. passargei – B. persica – B. petalidioides – B. petraea – B. phillipseae – B. pinguior – B. pratensis – B. procumbens – B. procurrens – B. pruinosa – B. pungens – B. pusilla – B. reekmansii – B. refracta – B. repens – B. richardsiae – B. rupicola – B. ruwenzoriensis – B. saturejaefolia – B. saxatilis – B. scandens – B. scindica – B. scullyi – B. sericea – B. serrulata – B. setosa – B. sindica – B. sinuata – B. sol – B. somaliensis – B. spathularis – B. spiculifolia – B. spinescens – B. spinifex – B. spinipes – B. squarrosa – B. stainbankiae – B. stuhlmanni – B. subglabra – B. subvolubilis – B. swaziensis – B. tanae – B. tanganyikensis – B. tanzaniensis – B. teaguei – B. tenuiramea – B. tetrasticha – B. thulinii – B. togodelia – B. torrei – B. transvaalensis – B. trifida – B. trinervis – B. trispina – B. trispinosa – B. turkanae – B. uniflora – B. uzondoensis – B. verdickii – B. verticillata – B. villosa – B. welwitschii – B. zollingeri
Name

Blepharis Juss., Gen. Pl. 103 (1789).

Type species: Blepharis boerhaviifolia Pers. (1807) (= Blepharis maderaspatensis Heyne ex Roth (1821).

Synonyms

Heterotypic
Acanthodium Delile, Descr. Egypte, Hist. Nat. 2(Mém.): 241 (1813).
Blepharacanthus Nees in J.Lindley, Intr. Nat. Syst. Bot., ed. 2: 444 (1836).
Trichacanthus Zoll. & Moritzi, Natuur- Geneesk. Arch. Ned.-Indië 2: 572 (1845).

References
Primary references

Jussieu, A.L. de 1789. Genera plantarum, secundum ordines naturales disposita juxta methodum in Horto Regio Parisiensi exaratam. 498 pp. Paris: Herissant et Theophile Barrois. BHL Reference page. 103

Additional references

Vollesen, K.B. 2000. Blepharis (Acanthaceae): A taxonomic revision. 342 pp., Royal Botanic Gardens, Kew. ISBN 1-900347-93-8. Reference page.

Links

Global Biodiversity Information Facility. 2019. GBIF Backbone Taxonomy. Checklist dataset. Taxon: Blepharis. .
International Plant Names Index. 2015. Blepharis. Published online. Accessed: Dec. 23 2015.
Tropicos.org 2015. Blepharis. Missouri Botanical Garden. Published online. Accessed: 23 Dec. 2015.
Hassler, M. 2021. World Plants. Synonymic Checklist and Distribution of the World Flora. . Blepharis. Accessed: 6 Jul 2021.
Hassler, M. 2021. Blepharis. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2021. Species 2000 & ITIS Catalogue of Life. Published online. Accessed: 2021 Jul 6. Reference page.
Govaerts, R. et al. 2021. Blepharis in Kew Science Plants of the World online. The Board of Trustees of the Royal Botanic Gardens, Kew. Published online. Accessed: 2021 Jul 6. Reference page.

Vernacular names
suomi: Pistiöt

Blepharis is a genus of plant in family Acanthaceae. It contains around 126 species found in seasonally dry to arid habitats from Africa over Arabia to Southeast Asia. In section Acanthodium, there are 13–15 species that use the C4 carbon fixation pathway. Phylogenetic analysis suggests that this pathway evolved up to three times independently in the genus over the last five million years.[1]
Species (selection)

Blepharis aequisepala Vollesen
Blepharis attenuata Napper
Blepharis boranensis Vollesen
Blepharis burundiensis Vollesen
Blepharis ciliaris
Blepharis crinita Juss.
Blepharis dhofarensis, A.G.Mill. [es; pt]
Blepharis diplodonta Vollesen
Blepharis drummondii Vollesen
Blepharis dunensis Vollesen
Blepharis duvigneaudii Vollesen
Blepharis edulis
Blepharis fenestralis Vollesen
Blepharis flava Vollesen
Blepharis gazensis Vollesen
Blepharis grossa T.Anderson
Blepharis gypsophila Thulin & Vollesen
Blepharis huillensis Vollesen
Blepharis inflata Vollesen
Blepharis inopinata Vollesen
Blepharis itigiensis Vollesen
Blepharis kenyensis Vollesen
Blepharis laevifolia Vollesen
Blepharis linariifolia Pers.
Blepharis macra (Nees) Vollesen
Blepharis maderaspatensis (L.) B.Heyne ex Roth
Blepharis meyeri Vollesen
Blepharis mitrata C.B.Clarke
Blepharis montana Vollesen
Blepharis obermeyerae Vollesen
Blepharis ogadenensis Vollesen
Blepharis petalidioides Vollesen
Blepharis petraea Vollesen
Blepharis pusilla Vollesen
Blepharis reekmansii Vollesen
Blepharis richardsiae Vollesen
Blepharis scandens Vollesen
Blepharis sericea Vollesen
Blepharis sindica
Blepharis somaliensis Vollesen
Blepharis spiculifolia Balf.f.
Blepharis spinescens Vollesen
Blepharis spinipes Vollesen
Blepharis subglabra Vollesen
Blepharis swaziensis Vollesen
Blepharis tanganyikensis (Napper) Vollesen
Blepharis tanzaniensis Vollesen
Blepharis thulinii Vollesen
Blepharis torrei Vollesen
Blepharis trifida Vollesen
Blepharis turkanae Vollesen
Blepharis uzondoensis Vollesen

References

Fisher, Amanda E.; McDade, Lucinda A.; Kiel, Carrie A.; Khoshravesh, Roxana; Johnson, Melissa A.; Stata, Matt; Sage, Tammy L.; Sage, Rowan F. (2015). "Evolutionary History of Blepharis (Acanthaceae) and the Origin of C4 Photosynthesis in Section Acanthodium". International Journal of Plant Sciences. 176 (8): 770–790. doi:10.1086/683011. ISSN 1058-5893. open access

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