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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Cladus: Core eudicots
Cladus: Asterids
Cladus: Lamiids
Ordo: Gentianales

Familia: Rubiaceae
Subfamilia: Rubioideae
Tribus: Coussareeae
Genera: Bradea – Coccocypselum – Coussarea – Cruckshanksia – Declieuxia – Faramea – Heterophyllaea – Hindsia – Oreopolus

Coussareeae Benth. & Hook.f., Gen. Pl. 2: 9, 24. (1873)

Type genus: Coussarea Aubl., Hist. Pl. Guiane 1: 98, t. 38. (1775)


Cruckshanksieae Benth. & Hook. f., 2: 9, 20. (1873)
Coccocypseleae Bremek., (1934)


Bentham, G. & Hooker, J.D. 1873. Genera Plantarum 2: 9, 24.
Bremer, B. & Eriksson, T. 2009. Time tree of Rubiaceae: phylogeny and dating the family, subfamilies, and tribes. International Journal of Plant Sciences 170(6): 766-793. DOI: 10.1086/599077 PDF Reference page.
Bremer, B. & Manen, J.F. 2000. Phylogeny and classification of the subfamily Rubioideae (Rubiaceae). Plant Systematics and Evolution 225(1-4): 43-72. DOI: 10.1007/BF00985458 PDF Reference page.
Delprete, P.G. & Jardim, J.G. 2012. Systematics, taxonomy and floristics of Brazilian Rubiaceae: an overview about the current status and future challenges. Rodriguésia 63(1): 101-128. DOI: 10.1590/S2175-78602012000100009 Reference page. 2018. Coussareeae. Missouri Botanical Garden. Published on the internet. Accessed: 2018 Dec. 17. Rubiaceae: Coussareeae. Missouri Botanical Garden. Published on the internet. Accessed: 2018 Dec. 17.

Coussareeae is a tribe of flowering plants in the family Rubiaceae and contains 407 species in 10 genera. The former tribe Coccocypseleae Bremek., consisting of Coccocypselum, Declieuxia, and Hindsia, is considered part of Coussareeae. Its representatives are found in Central and South America.[1]


Currently accepted names[1][2][3][4]

Bradea Standl. (6 sp)
Coccocypselum P.Browne (22 sp)
Coussarea Aubl. (120 sp)
Cruckshanksia Hook. & Arn. (7 sp)
Declieuxia Kunth (29 sp)
Faramea Aubl. (204 sp)
Heterophyllaea Hook.f. (2 sp)
Hindsia Benth. ex Lindl. (11 sp)
Oreopolus Schltdl. (1 sp)
Standleya Brade (6 sp)


Antoniana Tussac = Faramea
Bellardia Schreb. = Coccocypselum
Billardiera Vahl = Coussarea
Cococipsilum J.St.-Hil. = Coccocypselum
Condalia Ruiz & Pav. = Coccocypselum
Congdonia Jeps. = Declieuxia
Darluca Raf. = Faramea
Encopea C.Presl = Faramea
Evea Aubl. = Faramea
Froelichia Vahl = Coussarea
Homaloclados Hook.f. = Faramea
Lecanosperma Rusby = Heterophyllaea
Lipostoma D.Don = Coccocypselum
Macrosiphon Miq. = Hindsia
Neleixa Raf. = Faramea
Pecheya Scop. = Coussarea
Peckeya Raf. = Coussarea
Potima R.Hedw. = Faramea
Rotheria Meyen = Cruckshanksia
Sicelium P.Browne = Coccocypselum
Sulzeria Roem. & Schult. = Faramea
Tetramerium C.F.Gaertn. = Faramea
Thiersia Baill. = Faramea
Tontanea Aubl. = Coccocypselum


"World Checklist of Rubiaceae". Retrieved 14 April 2016.
Bremer B (2009). "A review of molecular phylogenetic studies of Rubiaceae". Annals of the Missouri Botanical Garden. 96: 4–26. doi:10.3417/2006197.
Bremer B, Eriksson E (2009). "Time tree of Rubiaceae: phylogeny and dating the family, subfamilies, and tribes". International Journal of Plant Sciences. 170: 766–793. doi:10.1086/599077.
Robbrecht E, Manen J-F (2006). "The major evolutionary lineages of the coffee family (Rubiaceae, angiosperms). Combined analysis (nDNA and cpDNA) to infer the position of Coptosapelta and Luculia, and supertree construction based on rbcL, rps16, trnL-trnF and atpB-rbcL data. A new classification in two subfamilies, Cinchonoideae and Rubioideae". Systematic Geography of Plants. 76: 85–146.

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