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Classification System: APG IV

Superregnum: Eukaryota
Regnum: Plantae
Cladus: Angiosperms
Cladus: Eudicots
Ordo: Ranunculales

Familia: Berberidaceae
Subfamilia: Podophylloideae
Genus: Diphylleia
Species: D. cymosaD. grayiD. sinensis

Diphylleia Michx., Fl. Bor.-Amer. 1: 203. t. 19-20. 19 Mar 1803.

Type species: D. cymosa

Michaux, A., 1803. Fl. Bor.-Amer. 1: 203. t. 19-20.
Farr, E. R. & Zijlstra, G. eds. (1996-) Index Nominum Genericorum (Plantarum). 2009 Oct 01 [1].
Ying T.S., S. Terabayashi, and D.E. Boufford. 1984. A monograph of Diphylleia (Berberidaceae). J. Arnold Arbor. 65: 57–94.


Hassler, M. 2019. Diphylleia. World Plants: Synonymic Checklists of the Vascular Plants of the World In: Roskovh, Y., Abucay, L., Orrell, T., Nicolson, D., Bailly, N., Kirk, P., Bourgoin, T., DeWalt, R.E., Decock, W., De Wever, A., Nieukerken, E. van, Zarucchi, J. & Penev, L., eds. 2019. Species 2000 & ITIS Catalogue of Life. Published on the internet. Accessed: 2019 Apr. 22. Reference page.
International Plant Names Index. 2019. Diphylleia. Published online. Accessed: Apr. 22 2019.
The Plant List 2013. Diphylleia in The Plant List Version 1.1. Published on the internet. Accessed: 2019 Apr. 22.
Tropicos.org 2019. Diphylleia. Missouri Botanical Garden. Published on the internet. Accessed: 2019 Apr. 22.

Vernacular names
suomi: Sontikat

Diphylleia is a group of small herbs in the family Berberidaceae described as a genus in 1803.[1][2] It is native to the eastern United States and eastern Asia.[3][4]

Diphylleia grayi, also known as the skeleton flower, has white petals that turn translucent with rain. When dry, they revert to white.[5]


Diphylleia cymosa Michx. - southern Appalachians from SW Virginia to NW Georgia
Diphylleia grayi F.Schmidt - Cape Sōya in northern Japan[7]
Diphylleia sinensis H.L.Li - China (Gansu, Hubei, Shaanxi, Sichuan, Yunnan)


Michaux, André. 1803. Flora Boreali-Americana 1: 203, plates 19–20. in Latin
Tropicos, Diphylleia Michx.
Flora of North America, Vol. 3 Diphylleia Michaux
Flora of China Vol. 19 Page 787 山荷叶属 shan he ye shu Diphylleia Michaux
J. Mater: Chem A, 2015, 3, 9379-9384
The Plant List, Kew Royal Botanic Gardens, Diphylleia Michx.
New York Botanical Garden Virtual Herbarium

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